Dear Authors,

Thank you for your thorough and detailed response to the feedback from two anonymous reviewers and one commenter. As you have undoubtedly noted, there is broad support for the work presented in the manuscript, but also a broad range of suggestions to better contextualize and better justify many of the claims made. These warrant a major revision, which your responses to the reviewers clearly show your author team is well positioned to undertake. I look forward to seeing the revisions you have foreshadowed in the response to the referees comments put into action.

Yours truly,

Sally Thompson

Dear Authors,

Thank you for your careful response to the detailed referee comments. It was clear that concerns were primarily related to matters of framing and inferring conclusions from the datasets, and that the data themselves and the study system were broadly of interest. Your clarifications, and willingness to temper some of the original claims of the study have clearly addressed the concerns that were present. I do not believe the reviewers need to see it again and I'm recommending publication at this point. Congratulations,

Sally Thompson