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  <front>
    <journal-meta><journal-id journal-id-type="publisher">HESS</journal-id><journal-title-group>
    <journal-title>Hydrology and Earth System Sciences</journal-title>
    <abbrev-journal-title abbrev-type="publisher">HESS</abbrev-journal-title><abbrev-journal-title abbrev-type="nlm-ta">Hydrol. Earth Syst. Sci.</abbrev-journal-title>
  </journal-title-group><issn pub-type="epub">1607-7938</issn><publisher>
    <publisher-name>Copernicus Publications</publisher-name>
    <publisher-loc>Göttingen, Germany</publisher-loc>
  </publisher></journal-meta>
    <article-meta>
      <article-id pub-id-type="doi">10.5194/hess-30-2879-2026</article-id><title-group><article-title>From soil to stream: modeling the catchment-scale hydrological effects of increased soil organic carbon</article-title><alt-title>From soil to stream</alt-title>
      </title-group>
      <contrib-group>
        <contrib contrib-type="author" corresp="yes" rid="aff1 aff2 aff3 aff5">
          <name><surname>Heinz</surname><given-names>Malve</given-names></name>
          <email>malve.heinz@unibe.ch</email>
        <ext-link>https://orcid.org/0000-0002-7020-2025</ext-link></contrib>
        <contrib contrib-type="author" corresp="no" rid="aff1 aff2">
          <name><surname>Holzkämper</surname><given-names>Annelie</given-names></name>
          
        <ext-link>https://orcid.org/0000-0002-1951-1041</ext-link></contrib>
        <contrib contrib-type="author" corresp="no" rid="aff4">
          <name><surname>Kumar</surname><given-names>Rohini</given-names></name>
          
        <ext-link>https://orcid.org/0000-0002-4396-2037</ext-link></contrib>
        <contrib contrib-type="author" corresp="no" rid="aff2">
          <name><surname>Ledain</surname><given-names>Sélène</given-names></name>
          
        </contrib>
        <contrib contrib-type="author" corresp="no" rid="aff1 aff3">
          <name><surname>Horton</surname><given-names>Pascal</given-names></name>
          
        <ext-link>https://orcid.org/0000-0003-0466-0359</ext-link></contrib>
        <contrib contrib-type="author" corresp="no" rid="aff1 aff3">
          <name><surname>Schaefli</surname><given-names>Bettina</given-names></name>
          
        <ext-link>https://orcid.org/0000-0003-1140-6244</ext-link></contrib>
        <aff id="aff1"><label>1</label><institution>Oeschger Centre for Climate Change Research, University of Bern, Bern, Switzerland</institution>
        </aff>
        <aff id="aff2"><label>2</label><institution>Agroecology and Environment, Agroscope, Zürich, Switzerland</institution>
        </aff>
        <aff id="aff3"><label>3</label><institution>Institute of Geography, University of Bern, Bern, Switzerland</institution>
        </aff>
        <aff id="aff4"><label>4</label><institution>Department of Computational Hydrosystems, Helmholtz Centre for Environmental Research – UFZ, Leipzig, Germany</institution>
        </aff>
        <aff id="aff5"><label>🏅</label><institution>Invited contribution by Malve Heinz, recipient of the EGU Hydrological Sciences Outstanding Student   and PhD candidate Presentation Award 2025.</institution>
        </aff>
      </contrib-group>
      <author-notes><corresp id="corr1">Malve Heinz (malve.heinz@unibe.ch)</corresp></author-notes><pub-date><day>13</day><month>May</month><year>2026</year></pub-date>
      
      <volume>30</volume>
      <issue>9</issue>
      <fpage>2879</fpage><lpage>2911</lpage>
      <history>
        <date date-type="received"><day>3</day><month>November</month><year>2025</year></date>
           <date date-type="rev-request"><day>21</day><month>November</month><year>2025</year></date>
           <date date-type="rev-recd"><day>17</day><month>April</month><year>2026</year></date>
           <date date-type="accepted"><day>21</day><month>April</month><year>2026</year></date>
      </history>
      <permissions>
        <copyright-statement>Copyright: © 2026 Malve Heinz et al.</copyright-statement>
        <copyright-year>2026</copyright-year>
      <license license-type="open-access"><license-p>This work is licensed under the Creative Commons Attribution 4.0 International License. To view a copy of this licence, visit <ext-link ext-link-type="uri" xlink:href="https://creativecommons.org/licenses/by/4.0/">https://creativecommons.org/licenses/by/4.0/</ext-link></license-p></license></permissions><self-uri xlink:href="https://hess.copernicus.org/articles/30/2879/2026/hess-30-2879-2026.html">This article is available from https://hess.copernicus.org/articles/30/2879/2026/hess-30-2879-2026.html</self-uri><self-uri xlink:href="https://hess.copernicus.org/articles/30/2879/2026/hess-30-2879-2026.pdf">The full text article is available as a PDF file from https://hess.copernicus.org/articles/30/2879/2026/hess-30-2879-2026.pdf</self-uri>
      <abstract><title>Abstract</title>

      <p id="d2e159">Droughts are increasingly threatening agricultural productivity. One potential adaptation is to increase the soil water retention capacity, which can be achieved by increasing soil organic carbon (SOC) through agricultural management. We investigated how increasing SOC affected catchment-scale hydrology including extremes. SOC increases were implemented via adjustments to soil hydraulic parameters (<inline-formula><mml:math id="M1" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi mathvariant="normal">b</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M2" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">PWP</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M3" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">FC</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M4" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">Sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M5" display="inline"><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>) in a mesoscale hydrologic modeling (mHM) framework, following literature-reported effects. Our analysis focuses on the medium-sized, agriculturally dominated Broye catchment in Western Switzerland and four nested subcatchments, where we evaluated five SOC increase scenarios of varying depth and magnitude. At the catchment scale, increment of SOC resulted in higher soil water content (1.43 %–3.75 %) and slightly higher evapotranspiration (0.15 %–0.38 %), while subsurface runoff was reduced (0.27 %–0.70 % across all scenarios). These values represent overall net changes over 2016–2022; while at shorter timescales, the magnitude and even direction of effects varied seasonally and by subcatchment. Increased water retention meant more soil water was available for evapotranspiration and less for groundwater recharge and streamflow. Consequently, streamflows were slightly reduced, peak flows modestly attenuated while low flow responses depended on catchment characteristics and timing. In warmer and drier subcatchments, low flow frequency increased in some years, whereas in cooler and wetter subcatchments, conditions in spring and early summer produced a beneficial effect, slightly reducing low flow frequency. Overall, our analysis suggests that large-scale increases in SOC can provide hydrological benefits such as enhanced agricultural productivity and reduced peak flows, but may involve trade-offs through reduced groundwater recharge and thus water availability.</p>
  </abstract>
    </article-meta>
  </front>
<body>
      

<sec id="Ch1.S1" sec-type="intro">
  <label>1</label><title>Introduction</title>
      <p id="d2e226">Agricultural productivity is strongly influenced by hydro-climatic variability. Meteorological, hydrological, and soil moisture droughts can co-occur and substantially reduce crop yields <xref ref-type="bibr" rid="bib1.bibx119 bib1.bibx51" id="paren.1"/>. Compared to soils under natural vegetation, agricultural soils are more prone to soil moisture depletion <xref ref-type="bibr" rid="bib1.bibx132" id="paren.2"/>. This vulnerability is also a consequence of long-term soil degradation: i.e., intensive management has depleted soil organic carbon <xref ref-type="bibr" rid="bib1.bibx113 bib1.bibx26" id="paren.3"/>, heavy machinery has compacted soils, increased surface runoff and reduced hydraulic conductivity and water retention <xref ref-type="bibr" rid="bib1.bibx59" id="paren.4"/>, and bare fallow practices have promoted erosion and weakened soil structure <xref ref-type="bibr" rid="bib1.bibx88" id="paren.5"/>. As a result, degraded soils have a reduced capacity to buffer hydro-climatic extremes, thereby amplifying both flood and drought impacts <xref ref-type="bibr" rid="bib1.bibx94" id="paren.6"/>. Under climate change, soil moisture droughts in Europe are expected to expand in both area and duration <xref ref-type="bibr" rid="bib1.bibx97" id="paren.7"/>.</p>
      <p id="d2e251">Drought impacts on agriculture arise from both plant responses and management constraints. Under combined precipitation and soil moisture deficits and high evaporative demand, plants reduce transpiration, affecting growth and yield quantity and quality <xref ref-type="bibr" rid="bib1.bibx32" id="paren.8"/>. Irrigation is a common strategy to mitigate drought stress and yield loss, even in water-rich regions like Switzerland <xref ref-type="bibr" rid="bib1.bibx131 bib1.bibx6" id="paren.9"/>. However, irrigation increasingly competes with ecological flow requirements and other water users <xref ref-type="bibr" rid="bib1.bibx15" id="paren.10"/>. In Switzerland, withdrawals from rivers may be restricted during low flow periods to protect aquatic ecosystems <xref ref-type="bibr" rid="bib1.bibx49" id="paren.11"/>, making yield losses unavoidable. Climate projections suggest that summer low flows in lowland Swiss catchments, including the Broye, are likely to become more frequent and severe from mid-century onward under climate change <xref ref-type="bibr" rid="bib1.bibx81" id="paren.12"/>, while projections for larger Central European catchments show mixed trends <xref ref-type="bibr" rid="bib1.bibx74" id="paren.13"/>. Hence, irrigation restrictions will probably become more frequent in the future.</p>
      <p id="d2e273">In this context, adapting agricultural management to strengthen the soil's function as a hydrological buffer, particularly its water retention capacity, can increase the resilience of agricultural cropping systems to droughts <xref ref-type="bibr" rid="bib1.bibx51" id="paren.14"/>. This buffering function is expressed through a set of soil hydraulic properties that control how water is stored and transmitted.</p>
      <p id="d2e279">A key metric for soil water availability is plant available water capacity (PAWC), defined as the difference in volumetric soil moisture between field capacity (<inline-formula><mml:math id="M6" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">FC</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>) and the permanent wilting point (<inline-formula><mml:math id="M7" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">PWP</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>). PAWC represents the range of soil water potentially accessible to plants, while the actual plant available water (PAW) denotes the fraction of PAWC currently present in the soil. Other parameters that are key to assessing the soil's hydraulic behavior are bulk density (<inline-formula><mml:math id="M8" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi mathvariant="normal">b</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>) and saturated hydraulic conductivity (<inline-formula><mml:math id="M9" display="inline"><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>). <inline-formula><mml:math id="M10" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi mathvariant="normal">b</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> describes the dry mass per total soil volume (commonly in g cm<sup>−3</sup>), while <inline-formula><mml:math id="M12" display="inline"><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> describes the rate at which water flows through saturated soil (cm d<sup>−1</sup>).</p>
      <p id="d2e374">Agricultural management practices that enhance soil structure and increase organic matter, such as conservation tillage, organic amendments, or cover cropping, can modify these parameters, particularly <inline-formula><mml:math id="M14" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi mathvariant="normal">b</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M15" display="inline"><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, and ultimately PAWC <xref ref-type="bibr" rid="bib1.bibx64 bib1.bibx13 bib1.bibx9 bib1.bibx22 bib1.bibx10" id="paren.15"/>. Increasing soil organic carbon (SOC) generally promotes soil aggregation and porosity, leading to lower <inline-formula><mml:math id="M16" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi mathvariant="normal">b</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, improved infiltration, <inline-formula><mml:math id="M17" display="inline"><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> and water retention, although the magnitude and direction of these effects depend on soil texture and structure <xref ref-type="bibr" rid="bib1.bibx104" id="paren.16"/>. <xref ref-type="bibr" rid="bib1.bibx49" id="text.17"/> showed that increasing SOC in potato fields can reduce drought stress and yield losses for a case study in Switzerland.</p>
      <p id="d2e431">Beyond these effects, increment of SOC offers a co-benefit of contributing to negative CO<sub>2</sub> emissions through carbon sequestration, particularly in the subsoil, a process encouraged by international initiatives such as the “4 per mille” initiative <xref ref-type="bibr" rid="bib1.bibx78 bib1.bibx16" id="paren.18"/>, as well as national and cantonal policies in Switzerland <xref ref-type="bibr" rid="bib1.bibx17 bib1.bibx30 bib1.bibx11" id="paren.19"/>. While in this context, mostly cropland is targeted, management-driven SOC gains have also been documented in permanent grasslands such as meadows and pastures, although their potential to increase SOC content is less well constrained yet <xref ref-type="bibr" rid="bib1.bibx87 bib1.bibx43 bib1.bibx57 bib1.bibx128" id="paren.20"/>. Assuming that such adaptive management to increase SOC is scaled up and applied on a larger area, it raises the question of how these field-level interventions affect catchment-scale hydrological processes.</p>
      <p id="d2e452">Local changes in land use can influence hydrologic processes at the catchment scale <xref ref-type="bibr" rid="bib1.bibx85 bib1.bibx82" id="paren.21"/>. In recent years, the use of natural processes to manage water in the landscape, often referred to as nature-based solutions, has received increasing attention <xref ref-type="bibr" rid="bib1.bibx23 bib1.bibx124" id="paren.22"/>. These practices include structural measures e.g. wetland and floodplain restoration, afforestation, riparian buffer strips, and terracing <xref ref-type="bibr" rid="bib1.bibx89 bib1.bibx61 bib1.bibx29" id="paren.23"/>). The latter has been studied in both modeling and field-based studies, showing potential to enhance soil moisture and reduce erosion locally, though possibly limiting downstream water availability <xref ref-type="bibr" rid="bib1.bibx29" id="paren.24"/>. Nature based solutions also include targeted arable soil management <xref ref-type="bibr" rid="bib1.bibx124" id="paren.25"/>, such as conservational tillage and gully treatment, which can decrease flood peaks and increase flood rise times, as observed in a data-based case study <xref ref-type="bibr" rid="bib1.bibx89" id="paren.26"/>. Modeling studies also indicate that practices like no-tillage can reduce hydraulic conductivity, leading to higher runoff and peak flows <xref ref-type="bibr" rid="bib1.bibx80 bib1.bibx61" id="paren.27"/>. <xref ref-type="bibr" rid="bib1.bibx34" id="paren.28"/> used the mechanistic model Tethys-Chloris <xref ref-type="bibr" rid="bib1.bibx33" id="paren.29"/> to analyze grassland management effects (e.g., grazing, mowing, compaction) from plot to catchment scale. They showed that detectable catchment-scale impacts are often subtle, require large interventions or long observation periods. <xref ref-type="bibr" rid="bib1.bibx33" id="text.30"/> highlight that modeling is often the only feasible way to assess such effects, given data limitations and the need for comparable catchments with and without management adaptations. To our knowledge, the impacts of field-scale agricultural management practices aimed to enhance soil water retention on evaporation, groundwater recharge, and hydrological extremes have not yet been systematically explored.</p>
      <p id="d2e486">Consequently, to investigate how field-level increases in SOC affect catchment-scale hydrology, including low and peak flows, we adopt a model-based approach. This is necessary because long-term observational data capturing pre- and post-management conditions are not available. We hypothesize that increasing SOC, and thus soil water retention, alters the timing and partitioning of water fluxes – potentially mitigating low-flow conditions by sustaining soil moisture and discharge during dry periods, while modestly reducing peak flows through enhanced retention capacity.</p>
      <p id="d2e489">For this analysis, we use the distributed mesoscale hydrological model mHM <xref ref-type="bibr" rid="bib1.bibx95 bib1.bibx63" id="paren.31"/>, representing agricultural management as scenarios of varying SOC increases. Changes in SOC propagate through the model via adjusted soil hydraulic parameters (<inline-formula><mml:math id="M19" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi mathvariant="normal">b</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M20" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">PWP</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M21" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">FC</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M22" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">Sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M23" display="inline"><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>), reflecting observed SOC effects reported in the literature. Our case study is the lowland, mid-sized agricultural Broye catchment in Western Switzerland, which is prone to agricultural droughts, summer low flows and has a good data coverage. We chose mHM for this analysis, as it is a distributed, open-source model under active development with a growing user community (<uri>https://mhm-ufz.org</uri>, last access: 10 May 2026). The model has been successfully used to simulate not only discharge, but also the spatiotemporal dynamics of runoff, evapotranspiration, and soil moisture across diverse European catchments <xref ref-type="bibr" rid="bib1.bibx95 bib1.bibx63 bib1.bibx100" id="paren.32"/>. mHM has also been applied to generate soil moisture time series for drought analysis and serves as the basis for the German drought monitor <xref ref-type="bibr" rid="bib1.bibx116 bib1.bibx97 bib1.bibx12" id="paren.33"/>.</p>
</sec>
<sec id="Ch1.S2">
  <label>2</label><title>Methods</title>
      <p id="d2e568">Our analysis framework is based on the catchment-scale hydrological model mHM, duly calibrated and evaluated using observed discharge time series (Sect. <xref ref-type="sec" rid="Ch1.S3.SS1"/>). Based on the reviewed literature (Sect. <xref ref-type="sec" rid="Ch1.S2.SS1"/>), we implement the effects of a theoretical combination of agricultural management practices for several scenarios of soil organic carbon (SOC) increases (Sect. <xref ref-type="sec" rid="Ch1.S2.SS7"/>). These scenarios are implemented by adjusting the input data for bulk density (<inline-formula><mml:math id="M24" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi mathvariant="normal">b</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>) using a pedotransfer function that considers SOC. The pedotransfer function used internally in mHM to calculate saturated hydraulic conductivity (<inline-formula><mml:math id="M25" display="inline"><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>) is also adjusted to consider SOC (see Sect. <xref ref-type="sec" rid="Ch1.S2.SS3"/>). We evaluate the effect of different SOC increase scenarios on the effective model parameters, hydrological states and fluxes at the grid scale and their effect on discharge including hydrological extremes.</p>
<sec id="Ch1.S2.SS1">
  <label>2.1</label><title>Literature-informed adjustment of soil hydraulic parameters</title>
      <p id="d2e609">We conducted a literature review to identify studies that examined changes in soil properties resulting from management adaptations aimed at increasing SOC (Table <xref ref-type="table" rid="T1"/>). Estimates of soil hydraulic properties – such as soil moisture at field capacity (<inline-formula><mml:math id="M26" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">FC</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>), permanent wilting point (<inline-formula><mml:math id="M27" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">PWP</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>), or saturated hydraulic conductivity (<inline-formula><mml:math id="M28" display="inline"><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>), derived from pedotransfer functions (PTFs) can vary considerably and are a source of uncertainty <xref ref-type="bibr" rid="bib1.bibx86 bib1.bibx122" id="paren.34"/>. However, since PTFs are (ideally) trained on large soil datasets from similar pedoclimatic conditions, they should support a broad generalization and enable the prediction of difficult-to-measure parameters from more easily observable ones. Moreover, they typically cover a wider range of soil textures than field or experimental studies.</p>

<table-wrap id="T1" specific-use="star"><label>Table 1</label><caption><p id="d2e653">Summary of soil property changes under different practices and modeling studies. PAWC = plant available water capacity (range between <inline-formula><mml:math id="M29" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">PWP</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> and <inline-formula><mml:math id="M30" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">FC</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>). SOM converted to SOC assuming SOM <inline-formula><mml:math id="M31" display="inline"><mml:mo>≈</mml:mo></mml:math></inline-formula> 58 % SOC.</p></caption><oasis:table frame="topbot"><oasis:tgroup cols="8">
     <oasis:colspec colnum="1" colname="col1" align="justify" colwidth="2cm"/>
     <oasis:colspec colnum="2" colname="col2" align="justify" colwidth="4.4cm"/>
     <oasis:colspec colnum="3" colname="col3" align="justify" colwidth="1.8cm"/>
     <oasis:colspec colnum="4" colname="col4" align="justify" colwidth="2cm"/>
     <oasis:colspec colnum="5" colname="col5" align="justify" colwidth="1.2cm"/>
     <oasis:colspec colnum="6" colname="col6" align="justify" colwidth="2cm"/>
     <oasis:colspec colnum="7" colname="col7" align="justify" colwidth="2.5cm"/>
     <oasis:colspec colnum="8" colname="col8" align="justify" colwidth="2.8cm"/>
     <oasis:thead>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1" align="left">Study type</oasis:entry>
         <oasis:entry colname="col2" align="left">Practice covered</oasis:entry>
         <oasis:entry colname="col3" align="left">Soil texture</oasis:entry>
         <oasis:entry colname="col4" align="left"><inline-formula><mml:math id="M32" display="inline"><mml:mi mathvariant="normal">Δ</mml:mi></mml:math></inline-formula> SOC</oasis:entry>
         <oasis:entry colname="col5" align="left"><inline-formula><mml:math id="M33" display="inline"><mml:mi mathvariant="normal">Δ</mml:mi></mml:math></inline-formula> <inline-formula><mml:math id="M34" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi mathvariant="normal">b</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col6" align="left"><inline-formula><mml:math id="M35" display="inline"><mml:mi mathvariant="normal">Δ</mml:mi></mml:math></inline-formula> <inline-formula><mml:math id="M36" display="inline"><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col7" align="left"><inline-formula><mml:math id="M37" display="inline"><mml:mi mathvariant="normal">Δ</mml:mi></mml:math></inline-formula> <inline-formula><mml:math id="M38" display="inline"><mml:mi mathvariant="italic">θ</mml:mi></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col8" align="left">Source</oasis:entry>
       </oasis:row>
     </oasis:thead>
     <oasis:tbody>
       <oasis:row>
         <oasis:entry colname="col1" align="left">experiment study or review</oasis:entry>
         <oasis:entry rowsep="1" colname="col2" align="left">Cover cropping</oasis:entry>
         <oasis:entry rowsep="1" colname="col3" align="left">various</oasis:entry>
         <oasis:entry rowsep="1" colname="col4" align="left"><inline-formula><mml:math id="M39" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>7 %–36 %</oasis:entry>
         <oasis:entry rowsep="1" colname="col5" align="left"><inline-formula><mml:math id="M40" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>1 % to <inline-formula><mml:math id="M41" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>4 %</oasis:entry>
         <oasis:entry rowsep="1" colname="col6" align="left"><inline-formula><mml:math id="M42" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>40 %–360 %</oasis:entry>
         <oasis:entry rowsep="1" colname="col7" align="left"><inline-formula><mml:math id="M43" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>4 %–20 % PAW</oasis:entry>
         <oasis:entry rowsep="1" colname="col8" align="left"><xref ref-type="bibr" rid="bib1.bibx46 bib1.bibx45" id="text.35"/></oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1" align="left"/>
         <oasis:entry rowsep="1" colname="col2" align="left">Organic farming (diverse crop rotations, organic amendments, manure application, tillage)</oasis:entry>
         <oasis:entry rowsep="1" colname="col3" align="left">various</oasis:entry>
         <oasis:entry rowsep="1" colname="col4" align="left"><inline-formula><mml:math id="M44" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>20 % to <inline-formula><mml:math id="M45" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>220 % (<sup>*</sup>1.2–<sup>*</sup>3.2)</oasis:entry>
         <oasis:entry rowsep="1" colname="col5" align="left"><inline-formula><mml:math id="M48" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>2 % to <inline-formula><mml:math id="M49" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>20 %</oasis:entry>
         <oasis:entry rowsep="1" colname="col6" align="left"><inline-formula><mml:math id="M50" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>50 %–250 %</oasis:entry>
         <oasis:entry rowsep="1" colname="col7" align="left"><inline-formula><mml:math id="M51" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>4 %–54 % PAW</oasis:entry>
         <oasis:entry rowsep="1" colname="col8" align="left"><xref ref-type="bibr" rid="bib1.bibx10" id="text.36"/></oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1" align="left"/>
         <oasis:entry rowsep="1" colname="col2" align="left">(Long-term) organic manure application</oasis:entry>
         <oasis:entry rowsep="1" colname="col3" align="left">Silt loam</oasis:entry>
         <oasis:entry rowsep="1" colname="col4" align="left"><inline-formula><mml:math id="M52" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>60 % (topsoil)</oasis:entry>
         <oasis:entry rowsep="1" colname="col5" align="left"><inline-formula><mml:math id="M53" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>14 % (topsoil)</oasis:entry>
         <oasis:entry rowsep="1" colname="col6" align="left">Likely <inline-formula><mml:math id="M54" display="inline"><mml:mo>↑</mml:mo></mml:math></inline-formula> but not significant</oasis:entry>
         <oasis:entry rowsep="1" colname="col7" align="left"><inline-formula><mml:math id="M55" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>7.8 %–9.7 % <inline-formula><mml:math id="M56" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry rowsep="1" colname="col8" align="left"><xref ref-type="bibr" rid="bib1.bibx104" id="text.37"/></oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1" align="left"/>
         <oasis:entry rowsep="1" colname="col2" align="left">Organic manure <inline-formula><mml:math id="M57" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula> biochar application</oasis:entry>
         <oasis:entry rowsep="1" colname="col3" align="left">Sandy loam</oasis:entry>
         <oasis:entry rowsep="1" colname="col4" align="left">Likely <inline-formula><mml:math id="M58" display="inline"><mml:mo>↑</mml:mo></mml:math></inline-formula></oasis:entry>
         <oasis:entry rowsep="1" colname="col5" align="left"><inline-formula><mml:math id="M59" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>2.6 % to <inline-formula><mml:math id="M60" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>4.8 %</oasis:entry>
         <oasis:entry rowsep="1" colname="col6" align="left"><inline-formula><mml:math id="M61" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>25 %</oasis:entry>
         <oasis:entry rowsep="1" colname="col7" align="left">Likely <inline-formula><mml:math id="M62" display="inline"><mml:mo>↑</mml:mo></mml:math></inline-formula></oasis:entry>
         <oasis:entry rowsep="1" colname="col8" align="left"><xref ref-type="bibr" rid="bib1.bibx125" id="text.38"/></oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1" align="left"/>
         <oasis:entry colname="col2" align="left">Review on crop residue effect</oasis:entry>
         <oasis:entry rowsep="1" colname="col3" align="left">Silt loam</oasis:entry>
         <oasis:entry rowsep="1" colname="col4" align="left"><inline-formula><mml:math id="M63" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>1 % (mass)</oasis:entry>
         <oasis:entry rowsep="1" colname="col5" align="left"><inline-formula><mml:math id="M64" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>3 %</oasis:entry>
         <oasis:entry rowsep="1" colname="col6" align="left"><inline-formula><mml:math id="M65" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>96 %</oasis:entry>
         <oasis:entry rowsep="1" colname="col7" align="left"><inline-formula><mml:math id="M66" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>33 % PAWC</oasis:entry>
         <oasis:entry rowsep="1" colname="col8" align="left"><xref ref-type="bibr" rid="bib1.bibx9" id="text.39"/></oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1" align="left"/>
         <oasis:entry rowsep="1" colname="col2" align="left"/>
         <oasis:entry rowsep="1" colname="col3" align="left">Clay loam</oasis:entry>
         <oasis:entry rowsep="1" colname="col4" align="left"><inline-formula><mml:math id="M67" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>0.65 % (mass)</oasis:entry>
         <oasis:entry rowsep="1" colname="col5" align="left"><inline-formula><mml:math id="M68" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>9 %</oasis:entry>
         <oasis:entry rowsep="1" colname="col6" align="left"><inline-formula><mml:math id="M69" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>90 %</oasis:entry>
         <oasis:entry rowsep="1" colname="col7" align="left"><inline-formula><mml:math id="M70" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>65 % PAWC</oasis:entry>
         <oasis:entry rowsep="1" colname="col8" align="left"/>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1" align="left"/>
         <oasis:entry colname="col2" align="left">Analysis of soil databases on SOC effects on soil hydraulic properties</oasis:entry>
         <oasis:entry colname="col3" align="left">Various textures (US)</oasis:entry>
         <oasis:entry colname="col4" align="left"><inline-formula><mml:math id="M71" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>0.6 % (mass)</oasis:entry>
         <oasis:entry colname="col5" align="left"/>
         <oasis:entry colname="col6" align="left"/>
         <oasis:entry colname="col7" align="left"><inline-formula><mml:math id="M72" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>1.5 %–1.7 % PAWC</oasis:entry>
         <oasis:entry colname="col8" align="left"><xref ref-type="bibr" rid="bib1.bibx70" id="text.40"/></oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1" align="left"/>
         <oasis:entry colname="col2" align="left"/>
         <oasis:entry colname="col3" align="left"/>
         <oasis:entry colname="col4" align="left"><inline-formula><mml:math id="M73" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>1 % (mass)</oasis:entry>
         <oasis:entry colname="col5" align="left"/>
         <oasis:entry colname="col6" align="left"/>
         <oasis:entry colname="col7" align="left"><inline-formula><mml:math id="M74" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>2 %–5 % PAWC</oasis:entry>
         <oasis:entry colname="col8" align="left"><xref ref-type="bibr" rid="bib1.bibx84" id="text.41"/></oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1" align="left"/>
         <oasis:entry colname="col2" align="left"/>
         <oasis:entry colname="col3" align="left"/>
         <oasis:entry colname="col4" align="left"><inline-formula><mml:math id="M75" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>1.2 % (mass)</oasis:entry>
         <oasis:entry colname="col5" align="left"/>
         <oasis:entry colname="col6" align="left"/>
         <oasis:entry colname="col7" align="left">up to <inline-formula><mml:math id="M76" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>50 % PAWC</oasis:entry>
         <oasis:entry colname="col8" align="left"><xref ref-type="bibr" rid="bib1.bibx52" id="text.42"/></oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1" align="left">modeling study</oasis:entry>
         <oasis:entry colname="col2" align="left">Analysis of effect of SOC on soil properties (high clay content reduces impact)</oasis:entry>
         <oasis:entry rowsep="1" colname="col3" align="left">Coarse (Sand 50 %)</oasis:entry>
         <oasis:entry rowsep="1" colname="col4" align="left">1 % vs. 3 %; 3 % vs. 5 %</oasis:entry>
         <oasis:entry rowsep="1" colname="col5" align="left"/>
         <oasis:entry rowsep="1" colname="col6" align="left"><inline-formula><mml:math id="M77" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>20 % to <inline-formula><mml:math id="M78" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>85 %, <inline-formula><mml:math id="M79" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>0 % to <inline-formula><mml:math id="M80" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>7 %</oasis:entry>
         <oasis:entry rowsep="1" colname="col7" align="left"/>
         <oasis:entry rowsep="1" colname="col8" align="left"><xref ref-type="bibr" rid="bib1.bibx93" id="text.43"/></oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1" align="left"/>
         <oasis:entry rowsep="1" colname="col2" align="left"/>
         <oasis:entry rowsep="1" colname="col3" align="left">Fine (Sand 20 %)</oasis:entry>
         <oasis:entry rowsep="1" colname="col4" align="left">1 % vs. 3 %; 3 % vs. 5 %</oasis:entry>
         <oasis:entry rowsep="1" colname="col5" align="left"/>
         <oasis:entry rowsep="1" colname="col6" align="left"><inline-formula><mml:math id="M81" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>14 % to <inline-formula><mml:math id="M82" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>17 %, <inline-formula><mml:math id="M83" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>0 % to <inline-formula><mml:math id="M84" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>25 %</oasis:entry>
         <oasis:entry rowsep="1" colname="col7" align="left"/>
         <oasis:entry rowsep="1" colname="col8" align="left"/>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1" align="left"/>
         <oasis:entry rowsep="1" colname="col2" align="left">Modeling impacts of varying <inline-formula><mml:math id="M85" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi mathvariant="normal">b</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> on soil hydraulic properties</oasis:entry>
         <oasis:entry rowsep="1" colname="col3" align="left">Clay loam</oasis:entry>
         <oasis:entry rowsep="1" colname="col4" align="left"/>
         <oasis:entry rowsep="1" colname="col5" align="left"><inline-formula><mml:math id="M86" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>10 %</oasis:entry>
         <oasis:entry rowsep="1" colname="col6" align="left"><inline-formula><mml:math id="M87" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>127 %</oasis:entry>
         <oasis:entry rowsep="1" colname="col7" align="left"><inline-formula><mml:math id="M88" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>7 % <inline-formula><mml:math id="M89" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M90" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>8 % <inline-formula><mml:math id="M91" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">res</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry rowsep="1" colname="col8" align="left"><xref ref-type="bibr" rid="bib1.bibx60" id="text.44"/></oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1" align="left"/>
         <oasis:entry colname="col2" align="left"/>
         <oasis:entry rowsep="1" colname="col3" align="left">Silt loam</oasis:entry>
         <oasis:entry rowsep="1" colname="col4" align="left"/>
         <oasis:entry rowsep="1" colname="col5" align="left"><inline-formula><mml:math id="M92" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>10 %</oasis:entry>
         <oasis:entry rowsep="1" colname="col6" align="left"><inline-formula><mml:math id="M93" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>114 %</oasis:entry>
         <oasis:entry rowsep="1" colname="col7" align="left"><inline-formula><mml:math id="M94" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>7 % <inline-formula><mml:math id="M95" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M96" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>8.7 % <inline-formula><mml:math id="M97" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">res</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry rowsep="1" colname="col8" align="left"/>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1" align="left"/>
         <oasis:entry rowsep="1" colname="col2" align="left"/>
         <oasis:entry rowsep="1" colname="col3" align="left">Sandy loam</oasis:entry>
         <oasis:entry rowsep="1" colname="col4" align="left"/>
         <oasis:entry rowsep="1" colname="col5" align="left"><inline-formula><mml:math id="M98" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>10 %</oasis:entry>
         <oasis:entry rowsep="1" colname="col6" align="left"><inline-formula><mml:math id="M99" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>86.4 %</oasis:entry>
         <oasis:entry rowsep="1" colname="col7" align="left"><inline-formula><mml:math id="M100" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>7 % <inline-formula><mml:math id="M101" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M102" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>7 % <inline-formula><mml:math id="M103" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">res</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry rowsep="1" colname="col8" align="left"/>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1" align="left"/>
         <oasis:entry colname="col2" align="left">Modeling land use change scenarios</oasis:entry>
         <oasis:entry colname="col3" align="left">various</oasis:entry>
         <oasis:entry colname="col4" align="left"/>
         <oasis:entry colname="col5" align="left"><inline-formula><mml:math id="M104" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>5 %, <inline-formula><mml:math id="M105" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>10 %, <inline-formula><mml:math id="M106" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>15 %</oasis:entry>
         <oasis:entry colname="col6" align="left"><inline-formula><mml:math id="M107" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>35 %, <inline-formula><mml:math id="M108" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>70 %, <inline-formula><mml:math id="M109" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>140 %</oasis:entry>
         <oasis:entry colname="col7" align="left"><inline-formula><mml:math id="M110" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>5 %, <inline-formula><mml:math id="M111" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>20 %, <inline-formula><mml:math id="M112" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>30 % PAWC</oasis:entry>
         <oasis:entry colname="col8" align="left"><xref ref-type="bibr" rid="bib1.bibx13" id="text.45"/></oasis:entry>
       </oasis:row>
     </oasis:tbody>
   </oasis:tgroup></oasis:table></table-wrap>

      <p id="d2e1693">A range of management practices have been shown to increase SOC: including cover cropping, diversified crop rotations, the application of organic amendments (e.g., compost or manure), the retention of crop residues and the application of biochar (Table <xref ref-type="table" rid="T1"/>). These practices are often combined, and the magnitude of SOC increase varies depending on site-specific conditions, depth, and implementation duration. The reported increases in SOC range from 7 % to 36 %, 20 % to 220 %, and 60 % to absolute increases of approximately <inline-formula><mml:math id="M113" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>1 % by mass <xref ref-type="bibr" rid="bib1.bibx9 bib1.bibx104 bib1.bibx46 bib1.bibx45 bib1.bibx10" id="paren.46"/>.</p>
      <p id="d2e1709">In addition to changes in SOC, several studies report concurrent effects on other soil hydraulic properties. A reduction in bulk density (<inline-formula><mml:math id="M114" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi mathvariant="normal">b</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>) is frequently observed; the effect varies from <inline-formula><mml:math id="M115" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>1 % to <inline-formula><mml:math id="M116" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>4 % through cover cropping <xref ref-type="bibr" rid="bib1.bibx46 bib1.bibx45" id="paren.47"/> to <inline-formula><mml:math id="M117" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>14 % for a silt loam in response to long-term organic amendments <xref ref-type="bibr" rid="bib1.bibx104" id="paren.48"/>.</p>
      <p id="d2e1751">The ranges of change in saturated hydraulic conductivity (<inline-formula><mml:math id="M118" display="inline"><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>) are particularly variable, with reported increases of 50 % to 250 %, 40 % to 360 % and 95 % depending on the practice and the site. <inline-formula><mml:math id="M119" display="inline"><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> underlies large variability and is a generally hard-to-measure and even harder-to-estimate variable and should be handled with caution <xref ref-type="bibr" rid="bib1.bibx127" id="paren.49"/>.</p>
      <p id="d2e1779">Soil moisture (<inline-formula><mml:math id="M120" display="inline"><mml:mi mathvariant="italic">θ</mml:mi></mml:math></inline-formula>) and especially plant available water capacity (PAWC), are reported to increase in the range of 4 % to 20 %, 4 % to 54 %, 33 % and 65 %, following increasing SOC and decreasing <inline-formula><mml:math id="M121" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi mathvariant="normal">b</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> <xref ref-type="bibr" rid="bib1.bibx9 bib1.bibx46 bib1.bibx45" id="paren.50"/>.</p>
      <p id="d2e1803">In <xref ref-type="bibr" rid="bib1.bibx9" id="text.51"/>, reducing crop residue cover from 100 % to 0 % decreased SOC, increased <inline-formula><mml:math id="M122" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi mathvariant="normal">b</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, and reduced <inline-formula><mml:math id="M123" display="inline"><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> and PAWC. In Table <xref ref-type="table" rid="T1"/>, we assume that increasing residue cover from 0 % to 100 % would have the opposite effects: increasing SOC, reducing BD, and increasing <inline-formula><mml:math id="M124" display="inline"><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> and PAWC.</p>
      <p id="d2e1844">The role of tillage is more complex. While reduced or no-tillage is often associated with higher SOC in the topsoil, it primarily leads to a redistribution of organic matter, with less SOC in deeper layers, and total SOC differences are not always significant <xref ref-type="bibr" rid="bib1.bibx14" id="paren.52"/>. Tillage is often used in organic farming to control weeds, which can offset some of the beneficial effects of organic farming practices. In particular, <xref ref-type="bibr" rid="bib1.bibx10" id="text.53"/> describe how tillage can negatively affect bulk density (<inline-formula><mml:math id="M125" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi mathvariant="normal">b</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>) and <inline-formula><mml:math id="M126" display="inline"><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, potentially counteracting the positive impacts of increased SOC in organic management systems, depending on tillage frequency and intensity.</p>
</sec>
<sec id="Ch1.S2.SS2">
  <label>2.2</label><title>Model description</title>
      <p id="d2e1883">The mesoscale Hydrologic Model (mHM, v. 5.13.1; <uri>https://mhm-ufz.org</uri>, last access: 10 May 2026) is a spatially distributed, process-based model designed to simulate major hydrological processes and water balance across diverse hydroclimatic regions and scales <xref ref-type="bibr" rid="bib1.bibx95 bib1.bibx63 bib1.bibx39" id="paren.54"/>. The computation of soil moisture processes and the generation of mobile water takes place at a grid scale, followed by a HBV-like soil moisture-runoff transformation to transform grid-scale mobile water to grid-scale runoff, followed by transfer and routing from grid cell to grid cell following topography-based flow directions (see below). The multiscale-parameter regionalization (MPR) is a key feature of mHM, which allows for both high-resolution spatial input data and computational efficiency <xref ref-type="bibr" rid="bib1.bibx95 bib1.bibx63" id="paren.55"/>. Using transfer functions, effective model parameters (such as hydraulic conductivity) at the grid-scale are estimated from spatial input parameters such as soil texture. These effective parameters are then internally upscaled to the (coarser) model resolution using different operators such as harmonic or arithmetic mean, while retaining spatial variability <xref ref-type="bibr" rid="bib1.bibx95 bib1.bibx63" id="paren.56"/>. More detailed descriptions are available in the work of <xref ref-type="bibr" rid="bib1.bibx95 bib1.bibx63" id="text.57"/> with more specific details on soil hydraulic parameterizations in <xref ref-type="bibr" rid="bib1.bibx71" id="text.58"/>.</p>

      <fig id="F1" specific-use="star"><label>Figure 1</label><caption><p id="d2e1907">mHM flowdiagram adapted from <xref ref-type="bibr" rid="bib1.bibx63" id="text.59"/>.</p></caption>
          <graphic xlink:href="https://hess.copernicus.org/articles/30/2879/2026/hess-30-2879-2026-f01.png"/>

        </fig>

      <p id="d2e1919">The main processes simulated in mHM are canopy interception, snow accumulation and melt, evapotranspiration, infiltration, soil moisture storage, surface runoff, lateral subsurface flow (called interflow in mHM), percolation, groundwater storage, baseflow and in-stream routing <xref ref-type="bibr" rid="bib1.bibx95" id="paren.60"/> (Fig. <xref ref-type="fig" rid="F1"/>). Snow accumulation is simulated with a simple temperature threshold; snowmelt is based on a degree-day method. In mHM, surface runoff can only occur on (nearly) impervious grid cells representing sealed areas such as streets or buildings. Potential runoff from excess water is assumed to re-infiltrate at the grid-scale and is, therefore, not simulated as a separate process in mHM. This is justified by the typically recommended grid resolution of 1 km to 50 km <xref ref-type="bibr" rid="bib1.bibx95" id="paren.61"/>.</p>
      <p id="d2e1931">The soil moisture and runoff generation schemes in mHM are conceptually based on the HBV model <xref ref-type="bibr" rid="bib1.bibx8" id="paren.62"/>, with some differences: mHM simulates soil moisture dynamics per soil layer (HBV usually has a single layer); the routine is described in more detail in Sect. <xref ref-type="sec" rid="Ch1.S2.SS3"/>. Once mobile water is generated per soil layer, the HBV conceptualization is used to transform grid-scale mobile water into grid-scale runoff. Each grid cell uses two subsurface storages fed with the sum of the mobile water from the soil moisture routine. The upper unsaturated  storage generates faster responding interflow and the other slower responding baseflow (Fig. <xref ref-type="fig" rid="F1"/>). Fast interflow occurs only if the water level in the storage zone exceeds a threshold; slow interflow is a permanent flux governed by the water level in the first bucket <xref ref-type="bibr" rid="bib1.bibx71" id="paren.63"/>. The remaining water level in this zone is the base for the percolation flux, encoded as a linear function of the water level. The percolation feeds the deeper saturated zone, supposed to emulate groundwater storage, where baseflow is again parameterized as a linear function of the water level <xref ref-type="bibr" rid="bib1.bibx95" id="paren.64"/>.</p>
      <p id="d2e1947">The total generated runoff (interflows and baseflow) from each grid cell is routed through the modelling domain by the multiscale Routing Model (mRM), a key component of the model <xref ref-type="bibr" rid="bib1.bibx117" id="paren.65"/>. Grid-scale runoff is transferred from cell to cell following topography-based flow direction and flow accumulation map. The routing algorithm applies the kinematic wave equation with spatially varying flow celerity parameterized by slope <xref ref-type="bibr" rid="bib1.bibx117" id="paren.66"/>. An adaptive time-stepping scheme is used to ensure numerical stability across resolutions. <xref ref-type="bibr" rid="bib1.bibx105" id="text.67"/> developed the subgrid catchment conservation (SCC) routine specifically for mHM as an alternative to the commonly used D8 algorithm <xref ref-type="bibr" rid="bib1.bibx83" id="paren.68"/>. This approach addresses the catchment size problem that arises when small catchments are simulated at coarse resolution, which can lead to over- or underestimation of catchment area and the resulting streamflow. For cells intersecting several subcatchments, SCC allows water to partition into different neighboring cells. Due to this study's relatively small catchment size, we also employ the SCC algorithm, which reduces biases in discharge between different subcatchments <xref ref-type="bibr" rid="bib1.bibx105" id="paren.69"/>.</p>
      <p id="d2e1965">In the configuration chosen for this study (see Sect. <xref ref-type="sec" rid="Ch1.S2.SS6"/>), the model has 47 (global) parameters that are calibrated based on observed streamflow (calibrated parameter values are shown in Supplement Sect. S4, Table S3). mHM has a built-in calibration algorithm based on a dynamically dimensioned search algorithm <xref ref-type="bibr" rid="bib1.bibx120" id="paren.70"/> for single objective parameter optimization. The users can choose between several performance criteria (<uri>https://mhm-ufz.org</uri>,  last access: 10 May 2026). The retained calibration options for the case studies at hand are further discussed in Sect. <xref ref-type="sec" rid="Ch1.S2.SS6"/>. The number of iterations is set to 2500, which has been successfully used to calibrate the mHM model in other studies <xref ref-type="bibr" rid="bib1.bibx62 bib1.bibx96 bib1.bibx106" id="paren.71"/>.</p>
</sec>
<sec id="Ch1.S2.SS3">
  <label>2.3</label><title>Parameterization of mHM soil moisture dynamics related to SOC changes</title>
      <p id="d2e1989">The mHM model represents root-zone soil moisture dynamics across multiple soil layers, with each layer corresponding to an individual soil water reservoir. The water balance within each reservoir is primarily controlled by incoming fluxes – snowmelt and rainfall in the uppermost layer, or percolation from the overlying soil layer in lower layers – and outgoing fluxes, including downward percolation and layer-specific evapotranspiration. Each soil layer has an upper soil water limit, represented by <inline-formula><mml:math id="M127" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, which acts as a threshold for storage capacity. <inline-formula><mml:math id="M128" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> is estimated using the PTF by <xref ref-type="bibr" rid="bib1.bibx133" id="text.72"/> (mHM default):

            <disp-formula id="Ch1.E1" content-type="numbered"><label>1</label><mml:math id="M129" display="block"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub><mml:mo>=</mml:mo><mml:msub><mml:mi>C</mml:mi><mml:mi mathvariant="normal">constant</mml:mi></mml:msub><mml:mo>+</mml:mo><mml:msub><mml:mi>C</mml:mi><mml:mi mathvariant="normal">clay</mml:mi></mml:msub><mml:mo>⋅</mml:mo><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi mathvariant="normal">clay</mml:mi></mml:msub><mml:mo>+</mml:mo><mml:msub><mml:mi>C</mml:mi><mml:mi mathvariant="normal">DB</mml:mi></mml:msub><mml:mo>⋅</mml:mo><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi mathvariant="normal">b</mml:mi></mml:msub></mml:mrow></mml:math></disp-formula>

          where <inline-formula><mml:math id="M130" display="inline"><mml:mrow><mml:msub><mml:mi>C</mml:mi><mml:mi mathvariant="normal">clay</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> is the clay content, and <inline-formula><mml:math id="M131" display="inline"><mml:mrow><mml:msub><mml:mi>C</mml:mi><mml:mi mathvariant="normal">constant</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M132" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi mathvariant="normal">clay</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, and <inline-formula><mml:math id="M133" display="inline"><mml:mrow><mml:msub><mml:mi>C</mml:mi><mml:mi mathvariant="normal">DB</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> are (global) parameters that are calibrated (Sect. S4, Table S3). At each time step, the current water content <inline-formula><mml:math id="M134" display="inline"><mml:mi mathvariant="italic">θ</mml:mi></mml:math></inline-formula> in each soil layer is compared to <inline-formula><mml:math id="M135" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>; if <inline-formula><mml:math id="M136" display="inline"><mml:mi mathvariant="italic">θ</mml:mi></mml:math></inline-formula> is below saturation, infiltration into the layer is allowed. A portion of the incoming water is retained in the current layer, while the remainder percolates into the next layer (see Equations in Appendix <xref ref-type="sec" rid="App1.Ch1.S2.SS1"/>). This also means that if <inline-formula><mml:math id="M137" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> (i.e., the soil's water retention capacity) increases, then for the same water input, less water infiltrates to deeper layers.</p>

      <fig id="F2"><label>Figure 2</label><caption><p id="d2e2149">Adjustment of bulk density (<inline-formula><mml:math id="M138" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi mathvariant="normal">b</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>) for organic matter (<inline-formula><mml:math id="M139" display="inline"><mml:mrow><mml:msub><mml:mi>p</mml:mi><mml:mi mathvariant="normal">OM</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>) in the default mHM routine, compared to the model version adapted for this study using spatially distributed SOC data.</p></caption>
          <graphic xlink:href="https://hess.copernicus.org/articles/30/2879/2026/hess-30-2879-2026-f02.png"/>

        </fig>

      <p id="d2e2180">In the default mHM setup, bulk density (<inline-formula><mml:math id="M140" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi mathvariant="normal">b</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>) is internally estimated from a user-defined mineral bulk density and modified using an organic matter parameter (<inline-formula><mml:math id="M141" display="inline"><mml:mrow><mml:msub><mml:mi>p</mml:mi><mml:mi mathvariant="normal">OM</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>), which can be fixed or calibrated but is spatially uniform (Fig. <xref ref-type="fig" rid="F2"/>). Saturated hydraulic conductivity (<inline-formula><mml:math id="M142" display="inline"><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>) in mHM is derived using the pedotransfer function (PTF) from <xref ref-type="bibr" rid="bib1.bibx24" id="text.73"/>, based on sand and clay content. We modify this parameterization to evaluate the effect of different SOC scenarios by directly linking SOC to <inline-formula><mml:math id="M143" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi mathvariant="normal">b</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> and <inline-formula><mml:math id="M144" display="inline"><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> (Fig. <xref ref-type="fig" rid="F2"/>). Specifically, we bypass the internal <inline-formula><mml:math id="M145" display="inline"><mml:mrow><mml:msub><mml:mi>p</mml:mi><mml:mi mathvariant="normal">OM</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> routine and instead input SOC-adjusted <inline-formula><mml:math id="M146" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi mathvariant="normal">b</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> values directly, using the PTF from <xref ref-type="bibr" rid="bib1.bibx73" id="text.74"/>, adapted by <xref ref-type="bibr" rid="bib1.bibx31" id="text.75"/>:

            <disp-formula id="Ch1.E2" content-type="numbered"><label>2</label><mml:math id="M147" display="block"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi mathvariant="normal">b</mml:mi></mml:msub><mml:mo>=</mml:mo><mml:mn mathvariant="normal">1.660</mml:mn><mml:mo>-</mml:mo><mml:mn mathvariant="normal">0.318</mml:mn><mml:msqrt><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi mathvariant="normal">SOC</mml:mi></mml:msub></mml:mrow></mml:msqrt><mml:mo>,</mml:mo></mml:mrow></mml:math></disp-formula>

          where <inline-formula><mml:math id="M148" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi mathvariant="normal">SOC</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> is the SOC content. Here we follow <xref ref-type="bibr" rid="bib1.bibx77" id="text.76"/> who showed that SOC consistently affects <inline-formula><mml:math id="M149" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi mathvariant="normal">b</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> in a largely texture-independent way. This PTF was trained on the extensive USDA soil database and is therefore assumed to be transferable to our study region. By representing SOC changes through <inline-formula><mml:math id="M150" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi mathvariant="normal">b</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> in pedotransfer functions, the resulting soil hydraulic parameters naturally reflect SOC effects <xref ref-type="bibr" rid="bib1.bibx133" id="paren.77"/>.</p>
      <p id="d2e2344">Above mentioned adaptation also allows us to capture the observed relationship between increasing SOC and decreasing <inline-formula><mml:math id="M151" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi mathvariant="normal">b</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, which is generally linked to higher <inline-formula><mml:math id="M152" display="inline"><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> <xref ref-type="bibr" rid="bib1.bibx102" id="paren.78"/>. To incorporate the effect of increasing SOC onto <inline-formula><mml:math id="M153" display="inline"><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, we also replace the default PTF with the one proposed by <xref ref-type="bibr" rid="bib1.bibx126" id="text.79"/>, as listed in <xref ref-type="bibr" rid="bib1.bibx67" id="text.80"/>, which includes SOC and <inline-formula><mml:math id="M154" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi mathvariant="normal">b</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> as predictors:

            <disp-formula id="Ch1.E3" content-type="numbered"><label>3</label><mml:math id="M155" display="block"><mml:mtable rowspacing="0.2ex" class="split" displaystyle="true" columnalign="right left"><mml:mtr><mml:mtd><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:mtd><mml:mtd><mml:mrow><mml:mo>=</mml:mo><mml:msub><mml:mi>C</mml:mi><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mrow><mml:mi mathvariant="normal">sat</mml:mi><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msub></mml:mrow></mml:msub><mml:mi>exp⁡</mml:mi><mml:mo mathsize="1.5em">(</mml:mo><mml:msub><mml:mi>C</mml:mi><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mrow><mml:mi mathvariant="normal">sat</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:mrow></mml:msub></mml:mrow></mml:msub><mml:mo>-</mml:mo><mml:msub><mml:mi>C</mml:mi><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mrow><mml:mi mathvariant="normal">sat</mml:mi><mml:mn mathvariant="normal">3</mml:mn></mml:mrow></mml:msub></mml:mrow></mml:msub><mml:mi>ln⁡</mml:mi><mml:mo>(</mml:mo><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi mathvariant="normal">clay</mml:mi></mml:msub><mml:mo>)</mml:mo></mml:mrow></mml:mtd></mml:mtr><mml:mtr><mml:mtd/><mml:mtd><mml:mrow><mml:mo>-</mml:mo><mml:msub><mml:mi>C</mml:mi><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mrow><mml:mi mathvariant="normal">sat</mml:mi><mml:mn mathvariant="normal">4</mml:mn></mml:mrow></mml:msub></mml:mrow></mml:msub><mml:mi>ln⁡</mml:mi><mml:mo>(</mml:mo><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi mathvariant="normal">sand</mml:mi></mml:msub><mml:mo>)</mml:mo><mml:mo>-</mml:mo><mml:msub><mml:mi>C</mml:mi><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mrow><mml:mi mathvariant="normal">sat</mml:mi><mml:mn mathvariant="normal">5</mml:mn></mml:mrow></mml:msub></mml:mrow></mml:msub><mml:mi>ln⁡</mml:mi><mml:mo>(</mml:mo><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi mathvariant="normal">SOC</mml:mi></mml:msub><mml:mo>)</mml:mo><mml:mo>-</mml:mo><mml:msub><mml:mi>C</mml:mi><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mrow><mml:mi mathvariant="normal">sat</mml:mi><mml:mn mathvariant="normal">6</mml:mn></mml:mrow></mml:msub></mml:mrow></mml:msub><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi mathvariant="normal">b</mml:mi></mml:msub><mml:mo mathsize="1.5em">)</mml:mo><mml:mo>,</mml:mo></mml:mrow></mml:mtd></mml:mtr></mml:mtable></mml:math></disp-formula>

          where <inline-formula><mml:math id="M156" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi mathvariant="normal">clay</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> and <inline-formula><mml:math id="M157" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi mathvariant="normal">sand</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> are the clay and the sand content, the parameters <inline-formula><mml:math id="M158" display="inline"><mml:mrow><mml:msub><mml:mi>C</mml:mi><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mrow><mml:mi mathvariant="normal">sat</mml:mi><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msub></mml:mrow></mml:msub></mml:mrow></mml:math></inline-formula> to <inline-formula><mml:math id="M159" display="inline"><mml:mrow><mml:msub><mml:mi>C</mml:mi><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mrow><mml:mi mathvariant="normal">sat</mml:mi><mml:mn mathvariant="normal">6</mml:mn></mml:mrow></mml:msub></mml:mrow></mml:msub></mml:mrow></mml:math></inline-formula> are constants (values listed in Sect. S4, Table S2). This PTF was trained on a Belgian database which includes soils present in our study region. In mHM, <inline-formula><mml:math id="M160" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">FC</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> is parameterized as a function of <inline-formula><mml:math id="M161" display="inline"><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, such that higher <inline-formula><mml:math id="M162" display="inline"><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> corresponds to lower <inline-formula><mml:math id="M163" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">FC</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, based on the PTF derived from soil database analysis by <xref ref-type="bibr" rid="bib1.bibx123" id="text.81"/>.</p>
      <p id="d2e2656">These parameter adjustments propagate through the process simulation chain, and their effects on parameters, variables, states, and fluxes in response to increased SOC will be described and illustrated in the results Section (Fig. <xref ref-type="fig" rid="F6"/>). They influence the estimation of the van Genuchten parameters used to compute <inline-formula><mml:math id="M164" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M165" display="inline"><mml:mi mathvariant="italic">α</mml:mi></mml:math></inline-formula>, <inline-formula><mml:math id="M166" display="inline"><mml:mi>n</mml:mi></mml:math></inline-formula>, and <inline-formula><mml:math id="M167" display="inline"><mml:mi>m</mml:mi></mml:math></inline-formula>, as well as field capacity (<inline-formula><mml:math id="M168" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">FC</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>) and the permanent wilting point (<inline-formula><mml:math id="M169" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">PWP</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>) (Equations listed in Appendix <xref ref-type="sec" rid="App1.Ch1.S2.SS1"/>). These, in turn, affect the simulated soil moisture (<inline-formula><mml:math id="M170" display="inline"><mml:mi mathvariant="italic">θ</mml:mi></mml:math></inline-formula>) and the associated fluxes, including infiltration, evapotranspiration (ET), lateral subsurface flow, and percolation.</p>
      <p id="d2e2725">ET in mHM is computed as a reduction of potential evapotranspiration (PET) by a soil moisture stress factor, following the formulation of <xref ref-type="bibr" rid="bib1.bibx35" id="text.82"/> or <xref ref-type="bibr" rid="bib1.bibx56" id="text.83"/>. In this study, we used the mHM process representation of <xref ref-type="bibr" rid="bib1.bibx28" id="text.84"/>, which combines the Jarvis approach with a root distribution model based on <xref ref-type="bibr" rid="bib1.bibx53" id="text.85"/>. In this configuration, root density varies spatially and vertically as a function of soil field capacity (<inline-formula><mml:math id="M171" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">FC</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>).</p>
      <p id="d2e2752">The reduction from PET, after accounting for canopy interception, to ET is expressed as:

            <disp-formula id="Ch1.E4" content-type="numbered"><label>4</label><mml:math id="M172" display="block"><mml:mrow><mml:mi mathvariant="normal">ET</mml:mi><mml:mo>=</mml:mo><mml:mi mathvariant="normal">PET</mml:mi><mml:mo>⋅</mml:mo><mml:mi>f</mml:mi></mml:mrow></mml:math></disp-formula>

          where <inline-formula><mml:math id="M173" display="inline"><mml:mi>f</mml:mi></mml:math></inline-formula> is a soil moisture stress function defined by:

            <disp-formula id="Ch1.E5" content-type="numbered"><label>5</label><mml:math id="M174" display="block"><mml:mrow><mml:mi>f</mml:mi><mml:mo>=</mml:mo><mml:mfenced open="{" close=""><mml:mtable rowspacing="0.2ex" class="cases" columnspacing="1em" columnalign="left left" framespacing="0em"><mml:mtr><mml:mtd><mml:mrow><mml:mi>R</mml:mi><mml:mo>,</mml:mo></mml:mrow></mml:mtd><mml:mtd><mml:mrow><mml:mover accent="true"><mml:mi mathvariant="italic">θ</mml:mi><mml:mo mathvariant="normal">‾</mml:mo></mml:mover><mml:mo>≥</mml:mo><mml:msub><mml:mi>t</mml:mi><mml:mi mathvariant="normal">jarvis</mml:mi></mml:msub></mml:mrow></mml:mtd></mml:mtr><mml:mtr><mml:mtd><mml:mrow><mml:mi>R</mml:mi><mml:mfenced open="(" close=")"><mml:mstyle displaystyle="true"><mml:mfrac style="display"><mml:mover accent="true"><mml:mi mathvariant="italic">θ</mml:mi><mml:mo mathvariant="normal">‾</mml:mo></mml:mover><mml:mrow><mml:msub><mml:mi>t</mml:mi><mml:mi mathvariant="normal">jarvis</mml:mi></mml:msub></mml:mrow></mml:mfrac></mml:mstyle></mml:mfenced><mml:mo>,</mml:mo></mml:mrow></mml:mtd><mml:mtd><mml:mrow><mml:mover accent="true"><mml:mi mathvariant="italic">θ</mml:mi><mml:mo mathvariant="normal">‾</mml:mo></mml:mover><mml:mo>&lt;</mml:mo><mml:msub><mml:mi>t</mml:mi><mml:mi mathvariant="normal">jarvis</mml:mi></mml:msub></mml:mrow></mml:mtd></mml:mtr></mml:mtable></mml:mfenced></mml:mrow></mml:math></disp-formula></p>
      <p id="d2e2845">Here, <inline-formula><mml:math id="M175" display="inline"><mml:mrow><mml:msub><mml:mi>t</mml:mi><mml:mi mathvariant="normal">jarvis</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> is a calibrated threshold parameter, <inline-formula><mml:math id="M176" display="inline"><mml:mover accent="true"><mml:mi mathvariant="italic">θ</mml:mi><mml:mo mathvariant="normal">‾</mml:mo></mml:mover></mml:math></inline-formula> is the normalized soil water content:

            <disp-formula id="Ch1.E6" content-type="numbered"><label>6</label><mml:math id="M177" display="block"><mml:mrow><mml:mover accent="true"><mml:mi mathvariant="italic">θ</mml:mi><mml:mo mathvariant="normal">‾</mml:mo></mml:mover><mml:mo>=</mml:mo><mml:mstyle displaystyle="true"><mml:mfrac style="display"><mml:mrow><mml:mi mathvariant="italic">θ</mml:mi><mml:mo>-</mml:mo><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">pwp</mml:mi></mml:msub></mml:mrow><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub><mml:mo>-</mml:mo><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">pwp</mml:mi></mml:msub></mml:mrow></mml:mfrac></mml:mstyle><mml:mo>,</mml:mo></mml:mrow></mml:math></disp-formula>

          and <inline-formula><mml:math id="M178" display="inline"><mml:mi>R</mml:mi></mml:math></inline-formula> is the fraction of roots in each soil layer:

            <disp-formula id="Ch1.E7" content-type="numbered"><label>7</label><mml:math id="M179" display="block"><mml:mrow><mml:mi>R</mml:mi><mml:mo>=</mml:mo><mml:mfenced open="(" close=")"><mml:mrow><mml:mn mathvariant="normal">1</mml:mn><mml:mo>-</mml:mo><mml:msup><mml:msub><mml:mi>R</mml:mi><mml:mi mathvariant="normal">CoeffFC</mml:mi></mml:msub><mml:mrow><mml:msub><mml:mi>d</mml:mi><mml:mi mathvariant="normal">u</mml:mi></mml:msub></mml:mrow></mml:msup></mml:mrow></mml:mfenced><mml:mo>-</mml:mo><mml:mfenced close=")" open="("><mml:mrow><mml:mn mathvariant="normal">1</mml:mn><mml:mo>-</mml:mo><mml:msup><mml:msub><mml:mi>R</mml:mi><mml:mi mathvariant="normal">CoeffFC</mml:mi></mml:msub><mml:mrow><mml:msub><mml:mi>d</mml:mi><mml:mi mathvariant="normal">l</mml:mi></mml:msub></mml:mrow></mml:msup></mml:mrow></mml:mfenced></mml:mrow></mml:math></disp-formula>

          with <inline-formula><mml:math id="M180" display="inline"><mml:mrow><mml:msub><mml:mi>R</mml:mi><mml:mi mathvariant="normal">CoeffFC</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> representing the root fraction coefficient for the layer, and <inline-formula><mml:math id="M181" display="inline"><mml:mrow><mml:msub><mml:mi>d</mml:mi><mml:mi mathvariant="normal">u</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> and <inline-formula><mml:math id="M182" display="inline"><mml:mrow><mml:msub><mml:mi>d</mml:mi><mml:mi mathvariant="normal">l</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> denoting the upper and lower soil layer boundaries (Appendix <xref ref-type="sec" rid="App1.Ch1.S2.SS2"/>). This formulation allows soil-layer specific root fractions to modulate ET in response to soil moisture.</p>
</sec>
<sec id="Ch1.S2.SS4">
  <label>2.4</label><title>Study area</title>
      <p id="d2e3009">We apply the mHM model to the mid-sized (602 km<sup>2</sup>), lowland, pluvial Broye catchment in Western Switzerland (Fig. <xref ref-type="fig" rid="F3"/>). The modeling period is constrained by the availability of leaf area index input data and is therefore set to 2015–2022, with 2015 used as a warm-up period and discarded from the analysis. Despite the relatively short study period, there is considerable variability, with 2018 and 2022 as hot and dry years, 2017 as dry, 2016 and 2021 as cool and wet years, and some intermediate years (2019 and 2020, Fig. <xref ref-type="fig" rid="F3"/>).</p>

      <fig id="F3" specific-use="star"><label>Figure 3</label><caption><p id="d2e3027"><bold>(a)</bold> Subcatchments and gauging station locations and landcover classes. <bold>(b)</bold> Soil texture and soil organic carbon for the total Broye catchment. <bold>(c)</bold> Cumulative temperature and precipitation sums for the whole catchment, cumulative sum of discharge for subcatchments (scaled for easier comparison, in 2022 only data available for the Broye subcatchment).</p></caption>
          <graphic xlink:href="https://hess.copernicus.org/articles/30/2879/2026/hess-30-2879-2026-f03.png"/>

        </fig>

      <p id="d2e3044">The mHM model is set up for the entire Broye catchment domain, but discharge observations are available only for four nested subcatchments. One of these subcatchments is also named Broye and drains the largest area, with its outlet near the city of Payerne (Fig. <xref ref-type="fig" rid="F3"/>). For clarity, we refer to the full modeled domain as the Broye catchment and to the gauged subcatchments as Broye (subcatchment), Flon, Arbogne, and Petit Glâne.</p>
      <p id="d2e3050">The landscape is dominated by cropland interspersed with small forest patches. Soils are primarily loams, clay loams or sandy loams, with rather low SOC contents (averaging at 2.2 % in the topsoil, as shown in Fig. <xref ref-type="fig" rid="F3"/>). The region has a temperate climate, with mean annual precipitation of 1142 mm and mean annual temperature of 9.12 °C (1993–2022). The streams exhibit a typical pluvial flow regime, with discharge peaks in winter and low flows in summer, characteristic of lowland Swiss agricultural catchments.</p>
</sec>
<sec id="Ch1.S2.SS5">
  <label>2.5</label><title>Input data</title>
      <p id="d2e3063">The required input data and their sources are listed in Table <xref ref-type="table" rid="T2"/>. The morphological and land use input data have a resolution of 50 m <inline-formula><mml:math id="M184" display="inline"><mml:mo>×</mml:mo></mml:math></inline-formula> 50 m, and the meteorological data of 1 km <inline-formula><mml:math id="M185" display="inline"><mml:mo>×</mml:mo></mml:math></inline-formula> 1 km, which is also the internal modeling resolution. The water transfer and routing in the model are based on the provided flow direction. However, because the water flow in the flat part of the catchment is not well constrained by the DEM, a reconditioned DEM consistent with the mapped rivers must be calculated. After trying different tools that provided unsatisfactory results, we developed a new tool to seamlessly align DEMs with mapped stream networks, resulting in minimal terrain alteration: hydro-snap <xref ref-type="bibr" rid="bib1.bibx50" id="paren.86"/>. The approach is softer than a stream burn-in and alters the DEM only where necessary. It also constrains the flow direction to be consistent with a provided catchment boundary.</p>

<table-wrap id="T2" specific-use="star"><label>Table 2</label><caption><p id="d2e3088">Overview of input data used.</p></caption><oasis:table frame="topbot"><oasis:tgroup cols="2">
     <oasis:colspec colnum="1" colname="col1" align="justify" colwidth="3.5cm"/>
     <oasis:colspec colnum="2" colname="col2" align="justify" colwidth="10cm"/>
     <oasis:thead>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1" align="left">mHM input data</oasis:entry>
         <oasis:entry colname="col2" align="left">Data description and source</oasis:entry>
       </oasis:row>
     </oasis:thead>
     <oasis:tbody>
       <oasis:row rowsep="1">
         <oasis:entry namest="col1" nameend="col2" align="left">Morphological data with a 50 m resolution </oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1" align="left">Land use map</oasis:entry>
         <oasis:entry colname="col2" align="left">Land use reclassified in three classes: pervious, impervious, forest. Extracted from ESA WorldCover <xref ref-type="bibr" rid="bib1.bibx134" id="paren.87"/>.</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1" align="left">Soil map</oasis:entry>
         <oasis:entry colname="col2" align="left">Soil type map along with the corresponding table of soil horizons (texture %, bulk density g cm<sup>−3</sup>) <xref ref-type="bibr" rid="bib1.bibx110" id="paren.88"/>.</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1" align="left">Hydrogeological map</oasis:entry>
         <oasis:entry colname="col2" align="left">Map and corresponding table of the main hydrogeological classes <xref ref-type="bibr" rid="bib1.bibx36" id="paren.89"/>.</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1" align="left">Digital Elevevation Model</oasis:entry>
         <oasis:entry colname="col2" align="left">DEM reconditionned with hydro-snap <xref ref-type="bibr" rid="bib1.bibx50" id="paren.90"/> and based on the swissALTI3D product <xref ref-type="bibr" rid="bib1.bibx112" id="paren.91"/>.</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1" align="left">Flow direction map</oasis:entry>
         <oasis:entry colname="col2" align="left">Flow direction computed by pysheds <xref ref-type="bibr" rid="bib1.bibx5" id="paren.92"/> on the reconditionned DEM.</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1" align="left">Flow accumulation map</oasis:entry>
         <oasis:entry colname="col2" align="left">Flow accumulation computed from the flow direction map.</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1" align="left">Aspect map</oasis:entry>
         <oasis:entry colname="col2" align="left">Aspect map computed from the DEM</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1" align="left">Slope map</oasis:entry>
         <oasis:entry colname="col2" align="left">Slope map puted from the DEM</oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1" align="left">Gauges position map</oasis:entry>
         <oasis:entry colname="col2" align="left">Map with location of gauging stations</oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry namest="col1" nameend="col2" align="left">Forcing data with a 1000 m resolution </oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1" align="left">Precipitation</oasis:entry>
         <oasis:entry colname="col2" align="left">Daily precipitation (mm d<sup>−1</sup>) from the RhiresD dataset <xref ref-type="bibr" rid="bib1.bibx75" id="paren.93"/></oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1" align="left">Temperature</oasis:entry>
         <oasis:entry colname="col2" align="left">Average daily temperature (°C) from the TabsD dataset <xref ref-type="bibr" rid="bib1.bibx76" id="paren.94"/></oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1" align="left">PET</oasis:entry>
         <oasis:entry colname="col2" align="left">Daily PET calculated after Priestley-Taylor (mm d<sup>−1</sup>) using data from <xref ref-type="bibr" rid="bib1.bibx112 bib1.bibx76 bib1.bibx109" id="text.95"/></oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1" align="left">LAI</oasis:entry>
         <oasis:entry colname="col2" align="left">Monthly LAI derived from Sentinel 2 satellite data</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1" align="left">Discharge</oasis:entry>
         <oasis:entry colname="col2" align="left">Daily discharge (m<sup>3</sup> s<sup>−1</sup>) provided by <xref ref-type="bibr" rid="bib1.bibx37" id="text.96"/>, DGE-DIRNA-EAU (VD), personal communication, 2024; <xref ref-type="bibr" rid="bib1.bibx20" id="text.97"/>; <xref ref-type="bibr" rid="bib1.bibx21" id="text.98"/></oasis:entry>
       </oasis:row>
     </oasis:tbody>
   </oasis:tgroup></oasis:table></table-wrap>

      <p id="d2e3344">With the available gridded precipitation data <xref ref-type="bibr" rid="bib1.bibx75" id="paren.99"/>, the water balance in the subcatchments Petit Glâne and Arbogne does not close (Appendix <xref ref-type="sec" rid="App1.Ch1.S1"/>). The observed annual discharge is far too low compared to the catchment-average precipitation. However, comparable catchments nearby show similarly low discharge values <xref ref-type="bibr" rid="bib1.bibx18 bib1.bibx19" id="paren.100"/>; accordingly, discharge measurement errors alone cannot explain the difference. The gridded precipitation product we use might well contain interpolation artifacts given the substantial spatial variability of observed precipitation. Therefore, we also explored other precipitation products (Sect. S3). To reduce potential biases, we eventually combined the gridded precipitation product with data of the nearby meteo stations for the two smaller subcatchments, Arbogne and Petit Glâne (Sect. S3, Fig. S5).</p>
<sec id="Ch1.S2.SS5.SSS1">
  <label>2.5.1</label><title>LAI</title>
      <p id="d2e3363">It has been shown that using spatially distributed leaf area index (LAI) instead of monthly look-up tables improves the discharge estimation for the VIC model <xref ref-type="bibr" rid="bib1.bibx69" id="paren.101"/>, that mHM is partly based on <xref ref-type="bibr" rid="bib1.bibx115" id="paren.102"/>. Therefore, LAI was inferred from Sentinel-2 imagery using a specifically trained neural network (NN). Sentinel-2 provides multispectral data at up to 10 m resolution with a 3 d revisit time at mid-latitudes. To train the model, a radiative transfer model (PROSAIL; <xref ref-type="bibr" rid="bib1.bibx54" id="altparen.103"/>) was used to simulate vegetation spectral reflectances based on varying leaf and canopy parameters, thereby generating a training database. Here, PROSAIL was parametrised specifically for arable crops in Switzerland.</p>
      <p id="d2e3375">ESA’s Sentinel Application Platform (SNAP) toolbox includes a Biophysical Processor estimating LAI from Sentinel-2 imagery for all vegetation types <xref ref-type="bibr" rid="bib1.bibx130" id="paren.104"/>. We therefore used a two-model strategy: the generic SNAP model for forests, and a trained neural network for cropland. LAI was predicted at 10 m (NN) and 20 m (SNAP) resolution, then combined using our land-use mask (Tab.  <xref ref-type="table" rid="T2"/>, <xref ref-type="bibr" rid="bib1.bibx134" id="altparen.105"/>). Non-vegetated areas were set to zero. Monthly median values were calculated and upscaled to 50 m resolution using nearest-neighbor interpolation.</p>
</sec>
</sec>
<sec id="Ch1.S2.SS6">
  <label>2.6</label><title>Model set-up and evaluation</title>
      <p id="d2e3395">Different options are available to represent the hydrological processes in mHM (see <uri>https://mhm-ufz.org</uri>, last access: 10 May 2026, for details). We select the default options <xref ref-type="bibr" rid="bib1.bibx98" id="paren.106"/>, except for the soil moisture and the evapotranspiration routine. For the soil moisture routine, we select the option where ET in each soil layer is regulated by the relative available soil moisture, rather than being uniform across land use classes, implemented by <xref ref-type="bibr" rid="bib1.bibx28" id="text.107"/>. This option allows for a spatially varying root fraction distribution depending on the soil's field capacity (<inline-formula><mml:math id="M191" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">FC</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>), which is an advantage in the presence of a high-quality soil database of high resolution (90 m <inline-formula><mml:math id="M192" display="inline"><mml:mo>×</mml:mo></mml:math></inline-formula> 90 m)  <xref ref-type="bibr" rid="bib1.bibx110" id="paren.108"/>. In contrast to most crop and land surface models, where root distribution is prescribed as a depth-dependent function independent of soil moisture <xref ref-type="bibr" rid="bib1.bibx72" id="paren.109"/>, mHM explicitly links the root distribution to the soil's <inline-formula><mml:math id="M193" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">FC</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> following <xref ref-type="bibr" rid="bib1.bibx28" id="text.110"/>.</p>
      <p id="d2e3446">We compute PET as an external model input according to the Priestley-Taylor method <xref ref-type="bibr" rid="bib1.bibx90" id="paren.111"/>, which uses average temperature, solar radiation and elevation as input. We set the model options such that PET is further distributed in space based on aspect, as implemented by <xref ref-type="bibr" rid="bib1.bibx27" id="text.112"/>. It should be noted that the mHM option to correct PET based on LAI data led to unrealistically high PET/ET values in our case and was thus not used.</p>
      <p id="d2e3455">The model is calibrated using as performance criterion the Kling-Gupta efficiency (KGE) <xref ref-type="bibr" rid="bib1.bibx44" id="paren.113"/>, calculated on each of the observed streamflow time series (at the four gauges) and averaged thereafter (without weighting). The retained period for calibration is 2016–2019 and for evaluation 2020–2022. The specific calibration setting is the result of manual explorations of different objective functions and of number of iterations. With fitting the model based on the KGE, we could get the overall best performance while maintaining realistic dynamics of all states and fluxes. We also evaluate the model performance for soil moisture using observed timeseries of volumetric water content at three depths from a grassland site close to the gauging station of the Broye subcatchment in Payerne, measured as part of the Swiss Soil Moisture EXperiment SwissSMEX <xref ref-type="bibr" rid="bib1.bibx79" id="paren.114"/>. The model is run at a daily timestep.</p>
</sec>
<sec id="Ch1.S2.SS7">
  <label>2.7</label><title>SOC change experiments</title>
      <p id="d2e3472">We apply different scenarios to evaluate the effect of increased SOC on catchment hydrology to (i) represent possible outcomes from long-term agricultural management adaptations (Sect. <xref ref-type="sec" rid="Ch1.S2.SS1"/>), and (ii) test the model's sensitivity towards different levels and depths of SOC increases (Fig. <xref ref-type="fig" rid="F4"/>). In the Broye catchment, SOC is on average around 2.2 % in the first 30 cm (soil layer 1), and approximately 0.9 % between 30 and 60 cm (layer 2, Fig. <xref ref-type="fig" rid="F3"/>). The SOC ratio between layer 2 and layer 1 is therefore approximately 60 %. We apply this depth-decrease ratio to the increase scenarios 1, 3, and 5. These scenarios represent increasing magnitudes of SOC increases. In scenario 2, SOC is not increased in soil layer 2 at all, and in scenario 4, SOC is increased by 1 % (mass) in both layers.</p>

      <fig id="F4" specific-use="star"><label>Figure 4</label><caption><p id="d2e3483">SOC scenarios.</p></caption>
          <graphic xlink:href="https://hess.copernicus.org/articles/30/2879/2026/hess-30-2879-2026-f04.png"/>

        </fig>

      <p id="d2e3492">We emphasize that these scenarios are artificial and not intended to represent specific, immediately achievable management interventions, but should rather reflect the long-term possible outcomes of combinations of different management adaptations. While they are informed by the literature review (Sect. <xref ref-type="sec" rid="Ch1.S2.SS1"/>), the scenarios with large and deep SOC increases (MedC_MedC and vHighC_MedC) may be harder to achieve in practice. Nevertheless, including such scenarios allows us to explore the potential range of hydrological responses to SOC increases and test the sensitivity of the model to large changes in soil properties.</p>
      <p id="d2e3498">The SOC values in each scenario are then used to estimate the model input <inline-formula><mml:math id="M194" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi mathvariant="normal">b</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> (bulk density), as discussed in Sect. <xref ref-type="sec" rid="Ch1.S2.SS3"/>. Consistent with the rationale outlined in the introduction, we hypothesize that increasing SOC will enhance soil water retention, allowing the soil to buffer hydrologic extremes, reduce low-flow frequency, and modestly attenuate peak flows. mHM considers three land use types: forest, impervious cover and pervious cover, where the latter includes all cropland and meadows. The adaptations to <inline-formula><mml:math id="M195" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi mathvariant="normal">b</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> are only applied to pervious areas, which have the highest share in each subcatchment (shown in Fig. <xref ref-type="fig" rid="F3"/>, panel a). Within the pervious fraction, the actual land use composition differs across catchments: Petit Glâne and Arbogne are cropland-dominated (80 % and 66 % of pervious area), the Broye subcatchment is mixed (54 % cropland, 43 % permanent meadow), and the Flon is dominated by permanent meadows and pastures (80 % of pervious area).</p>
</sec>
<sec id="Ch1.S2.SS8">
  <label>2.8</label><title>Hydrologic extremes evaluations: Low and peak flow indicators</title>
      <p id="d2e3535">To assess the impact of the SOC scenarios on low flows, we calculate the <inline-formula><mml:math id="M196" display="inline"><mml:mrow><mml:msub><mml:mi>Q</mml:mi><mml:mn mathvariant="normal">347</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> threshold for each subcatchment. <inline-formula><mml:math id="M197" display="inline"><mml:mrow><mml:msub><mml:mi>Q</mml:mi><mml:mn mathvariant="normal">347</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> corresponds to the discharge that is exceeded on 347 d yr<sup>−1</sup> (i.e., the 5th percentile) and is commonly used in Switzerland to define low-flow periods and as a threshold for the restriction of irrigation water withdrawal from rivers <xref ref-type="bibr" rid="bib1.bibx111 bib1.bibx49" id="paren.115"/>.</p>
      <p id="d2e3575">For peak flows, our analysis is constrained by the daily resolution of simulated discharge, whereas hourly peaks would be more relevant <xref ref-type="bibr" rid="bib1.bibx4" id="paren.116"/>. Nevertheless, we estimate changes in discharge associated with two-year return period floods (<inline-formula><mml:math id="M199" display="inline"><mml:mrow><mml:msub><mml:mi>Q</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> events). <inline-formula><mml:math id="M200" display="inline"><mml:mrow><mml:msub><mml:mi>Q</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> thresholds are determined for each subcatchment using a generalized extreme value model, although the short time series in smaller subcatchments can be limiting (48 years in the Broye subcatchment and 26 years in the other subcatchments). Across stations, we also observe a decreasing discharge trend, significant only for the Broye subcatchment, which explains why only a few <inline-formula><mml:math id="M201" display="inline"><mml:mrow><mml:msub><mml:mi>Q</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> events occur during the study period.</p>
</sec>
</sec>
<sec id="Ch1.S3">
  <label>3</label><title>Results</title>
<sec id="Ch1.S3.SS1">
  <label>3.1</label><title>Calibration and evaluation</title>
      <p id="d2e3630">Model calibration led to a good fit of simulated to observed streamflow for the Broye subcatchment and the Flon (KGE <inline-formula><mml:math id="M202" display="inline"><mml:mo>=</mml:mo></mml:math></inline-formula> 0.91), with a slightly lower performance for the Arbogne and Petit Glâne (KGE <inline-formula><mml:math id="M203" display="inline"><mml:mo>=</mml:mo></mml:math></inline-formula> 0.83 and 0.86, Fig. <xref ref-type="fig" rid="F5"/>).</p>

      <fig id="F5" specific-use="star"><label>Figure 5</label><caption><p id="d2e3651">Observed and simulated discharge for all subcatchments. Except for the Broye subcatchment, the data for the other stations was not officially validated yet for 2022. NSE <inline-formula><mml:math id="M204" display="inline"><mml:mo>=</mml:mo></mml:math></inline-formula> Nash-Sutcliff-Efficiency, pbias <inline-formula><mml:math id="M205" display="inline"><mml:mo>=</mml:mo></mml:math></inline-formula> percentage bias.</p></caption>
          <graphic xlink:href="https://hess.copernicus.org/articles/30/2879/2026/hess-30-2879-2026-f05.png"/>

        </fig>

      <p id="d2e3674">Seasonal discharge dynamics are not equally well captured across subcatchments (Appendix <xref ref-type="sec" rid="App1.Ch1.S3.SS3"/> and <xref ref-type="sec" rid="App1.Ch1.S3.SS4"/>). The Broye subcatchment shows the best fit; low flows are underestimated in the Flon, overestimated and mis-timed in the Arbogne, and mis-timed in the Petit Glâne. Percentage biases for high and low flows (<inline-formula><mml:math id="M206" display="inline"><mml:mrow><mml:msub><mml:mi>Q</mml:mi><mml:mn mathvariant="normal">95</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> and <inline-formula><mml:math id="M207" display="inline"><mml:mrow><mml:msub><mml:mi>Q</mml:mi><mml:mn mathvariant="normal">5</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula>) range from <inline-formula><mml:math id="M208" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.1 % to 7 % and <inline-formula><mml:math id="M209" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>22 % to 43 %, respectively, with the best agreement in the Broye subcatchment (pbias for  <inline-formula><mml:math id="M210" display="inline"><mml:mrow><mml:msub><mml:mi>Q</mml:mi><mml:mn mathvariant="normal">95</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M211" display="inline"><mml:mo>=</mml:mo></mml:math></inline-formula> 7 %, pbias for <inline-formula><mml:math id="M212" display="inline"><mml:mrow><mml:msub><mml:mi>Q</mml:mi><mml:mn mathvariant="normal">5</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M213" display="inline"><mml:mo>=</mml:mo></mml:math></inline-formula> 9 %), likely reflecting the higher quality of observed discharge data there.</p>
      <p id="d2e3755">In comparison with the observed soil moisture time series (Sect. <xref ref-type="sec" rid="Ch1.S2.SS6"/>), mHM achieved reasonably good performance, except for the lowest soil layer, with KGE values of 0.65, 0.73, and 0.13 (0–30, 30–60 and 60–90 cm). The good fit in the two upper layers is noteworthy given that the data were not used for calibration and represent a single grid cell. While soil moisture was generally underestimated (percentage bias 8 % to <inline-formula><mml:math id="M214" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>11 %), the model reproduced the temporal variability well (Appendix <xref ref-type="sec" rid="App1.Ch1.S3.SS1"/>).</p>
</sec>
<sec id="Ch1.S3.SS2">
  <label>3.2</label><title>Change in soil hydraulic properties</title>
      <p id="d2e3777">In Fig. <xref ref-type="fig" rid="F6"/>a, we show the impact of the SOC increase on several parameters of interest. The points represent all pervious land cover cells in the overall catchment, which equals the area the SOC increase is applied to. Saturated hydraulic conductivity (<inline-formula><mml:math id="M215" display="inline"><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>) and water content at saturation and wilting point (<inline-formula><mml:math id="M216" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">Sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> and <inline-formula><mml:math id="M217" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">PWP</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>) are effective model parameters calculated within mHM. <inline-formula><mml:math id="M218" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">FC</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> changes with the almost same magnitude as <inline-formula><mml:math id="M219" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">Sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, which is why we do not show it explicitly in Fig. <xref ref-type="fig" rid="F6"/>. Bulk density (<inline-formula><mml:math id="M220" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi mathvariant="normal">b</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>) is a model input and PAWC (plant available water capacity, <inline-formula><mml:math id="M221" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">FC</mml:mi></mml:msub><mml:mo>-</mml:mo><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">PWP</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>) gives an idea if the surplus in retained water could be taken up by plants.</p>

      <fig id="F6" specific-use="star"><label>Figure 6</label><caption><p id="d2e3871"><bold>(a)</bold> Changes in key effective parameters for all pervious landcover cells, which represent the area where SOC was increased in our scenario runs. Please note the differences in scale for each plot. Number in each plot show the mean relative differences for each SOC scenario against the base scenario. <bold>(b)</bold> How changes in SOC and bulk density (<inline-formula><mml:math id="M222" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi mathvariant="normal">b</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>) propagate in the mHM model. The figure shows a snapshot of net changes in parameters and outputs; actual variables depend on boundary conditions, so seasonal responses may differ. Note that <inline-formula><mml:math id="M223" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">FC</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> scales linearly with <inline-formula><mml:math id="M224" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> but only weakly with <inline-formula><mml:math id="M225" display="inline"><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, leading to an overall increase. <inline-formula><mml:math id="M226" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="normal">f</mml:mi><mml:mi mathvariant="normal">runoff</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> and <inline-formula><mml:math id="M227" display="inline"><mml:mi>I</mml:mi></mml:math></inline-formula> at each timestep are controlled by the current <inline-formula><mml:math id="M228" display="inline"><mml:mi mathvariant="italic">θ</mml:mi></mml:math></inline-formula>; ET affects them only indirectly through its impact on <inline-formula><mml:math id="M229" display="inline"><mml:mi mathvariant="italic">θ</mml:mi></mml:math></inline-formula> in preceding timesteps, therefore the dashed arrow here.</p></caption>
          <graphic xlink:href="https://hess.copernicus.org/articles/30/2879/2026/hess-30-2879-2026-f06.png"/>

        </fig>

      <p id="d2e3962">Adding <inline-formula><mml:math id="M230" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>0.5 %, 1 % and 1.5 % SOC to the first soil layer led to a decrease in <inline-formula><mml:math id="M231" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi mathvariant="normal">b</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> by on average 3.3 %, 6.4 % and 9.2 % (Fig. <xref ref-type="fig" rid="F6"/>). The decrease in <inline-formula><mml:math id="M232" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi mathvariant="normal">b</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> propagates through the model (Fig. <xref ref-type="fig" rid="F1"/>), leading to averages increase by 3.2 %, 6.2 % and 8.8 % in both <inline-formula><mml:math id="M233" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">Sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> and <inline-formula><mml:math id="M234" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">FC</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>. PAWC is increased by 4.9 %, 9.3 % and up to 13.5 %. As described in Sect. <xref ref-type="sec" rid="Ch1.S2.SS3"/>, an increase in <inline-formula><mml:math id="M235" display="inline"><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> leads to a decrease in <inline-formula><mml:math id="M236" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">FC</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, as parameterized in mHM. Although <inline-formula><mml:math id="M237" display="inline"><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> was substantially increased, the effect on <inline-formula><mml:math id="M238" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">FC</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> is negligible. The sensitivity of PAWC to increases in SOC depends on the initial SOC content and soil texture. PAWC increases more strongly in soils with low initial SOC and higher sand content (Sect. S7).</p>
</sec>
<sec id="Ch1.S3.SS3">
  <label>3.3</label><title>Impact of increasing SOC on local and catchment-scale hydrological processes</title>
      <p id="d2e4077">We first isolate the grid cells where SOC-induced changes in soil hydraulic properties were applied (<inline-formula><mml:math id="M239" display="inline"><mml:mo lspace="0mm">=</mml:mo></mml:math></inline-formula> pervious landcover cells) to examine how these properties change and how states and fluxes respond locally. Subsequently, we aggregate the results to evaluate how these local changes propagate to influence hydrological states and fluxes at the catchment scale.</p>
      <p id="d2e4087">Figure <xref ref-type="fig" rid="F6"/>b illustrates how changes in hydraulic parameters propagate through the process chain in mHM and how states and fluxes are changed accordingly, here representing a snapshot for the simulated net changes. On average, increased soil water retention capacity leads to slightly higher ET and since more water can be retained and evaporated, less water contributes to further states and fluxes downwards.</p>
      <p id="d2e4092">The overall impact of the SOC scenarios at the grid-scale are moderate. Figure <xref ref-type="fig" rid="F7"/> shows actual evapotranspiration (ET), soil water content in the first and second soil layer, and subsurface runoff across all pervious land use cells in the overall catchment (other states and fluxes are shown in the Sect. S2). Subsurface runoff in mHM comprises the fluxes fast and slow interflow and baseflow. Figure <xref ref-type="fig" rid="F7"/> displays the range over all cells and the mean in solid lines: panel (a0 shows the timeseries, (b) shows the relative differences to the base scenario and panel (c) shows the cumulative differences. Average soil water content in layers 1 and 2 increases by average 2.9 % to 8.1 % (over all SOC scenarios), corresponding to 3–8 mm in winter and 2–6 mm in summer, with substantial spatial variability (Fig. <xref ref-type="fig" rid="F7"/>).  The impact of SOC increase on the boundary fluxes ET and subsurface is smaller: ET increases slightly (<inline-formula><mml:math id="M240" display="inline"><mml:mo lspace="0mm">+</mml:mo></mml:math></inline-formula>0.16 % to <inline-formula><mml:math id="M241" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>0.4 %) while subsurface runoff slightly decreases (<inline-formula><mml:math id="M242" display="inline"><mml:mi mathvariant="normal">−</mml:mi></mml:math></inline-formula>0.28 % to <inline-formula><mml:math id="M243" display="inline"><mml:mi mathvariant="normal">−</mml:mi></mml:math></inline-formula>0.72 %), corresponding to 0.2–0.6  and 1–2 mm, respectively. The differences in these key state and fluxes exhibit distinct seasonal patterns. For ET, differences peak in spring and summer, subsurface runoff peaks in winter are partly reverses in summer and fall. The difference in soil water content is largest in winter and spring, decreasing in late summer and early fall, before it sharply rising again in late fall.</p>

      <fig id="F7" specific-use="star"><label>Figure 7</label><caption><p id="d2e4133">All panels show the (weekly) mean and range over all pervious landcover cells, where SOC was increased. The legend applies to all panels. <bold>(a)</bold> Timeseries of key fluxes and state for the base and all SOC increase scenarios. Note, that the difference between the scenarios for ET and subsurface runoff is so small that the lines almost completely overlap. <bold>(b)</bold> Absolute difference between each SOC scenario and the base scenario. <bold>(c)</bold> Cumulative sums of the difference between each SOC scenario and the base scenario; the text in each subplot is the mean relative difference over all cells.</p></caption>
          <graphic xlink:href="https://hess.copernicus.org/articles/30/2879/2026/hess-30-2879-2026-f07.png"/>

        </fig>

      <fig id="F8"><label>Figure 8</label><caption><p id="d2e4153">Schematic of the annual cycle of average impacts of SOC increase scenarios relative to the base scenario.</p></caption>
          <graphic xlink:href="https://hess.copernicus.org/articles/30/2879/2026/hess-30-2879-2026-f08.png"/>

        </fig>

      <p id="d2e4162">We summarize the average seasonal pattern of all SOC scenarios relative to the base scenario in Fig. <xref ref-type="fig" rid="F8"/> and can distinguish four stages, defined in Table <xref ref-type="table" rid="T3"/>, which outline the main hydrological responses throughout the year. Overall, we see a consistent increase in soil moisture across all seasons, moderate increases in ET during spring and summer, and generally reduced subsurface runoff, except in spring when it shows a slight increase (Table <xref ref-type="table" rid="T3"/>).</p>

<table-wrap id="T3" specific-use="star"><label>Table 3</label><caption><p id="d2e4174">Mean annual hydrological impacts of increased SOC on key fluxes and states relative to base scenario.</p></caption><oasis:table frame="topbot"><oasis:tgroup cols="5">
     <oasis:colspec colnum="1" colname="col1" align="justify" colwidth="3.5cm"/>
     <oasis:colspec colnum="2" colname="col2" align="justify" colwidth="2cm"/>
     <oasis:colspec colnum="3" colname="col3" align="justify" colwidth="2cm"/>
     <oasis:colspec colnum="4" colname="col4" align="justify" colwidth="2cm"/>
     <oasis:colspec colnum="5" colname="col5" align="justify" colwidth="6.5cm"/>
     <oasis:thead>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1">Stage/Season</oasis:entry>
         <oasis:entry colname="col2">Soil Moisture</oasis:entry>
         <oasis:entry colname="col3">ET</oasis:entry>
         <oasis:entry colname="col4">Subsurface Runoff</oasis:entry>
         <oasis:entry colname="col5">Key Mechanism</oasis:entry>
       </oasis:row>
     </oasis:thead>
     <oasis:tbody>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1">Winter/early spring</oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M244" display="inline"><mml:mrow><mml:mo>↑</mml:mo><mml:mo>↑</mml:mo></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3">No difference</oasis:entry>
         <oasis:entry colname="col4"><inline-formula><mml:math id="M245" display="inline"><mml:mo>↓</mml:mo></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col5">High water retention capacity stores precipitation, minimizing subsurface runoff under SOC scenario.</oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1">Late spring/early summer</oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M246" display="inline"><mml:mo>↑</mml:mo></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3"><inline-formula><mml:math id="M247" display="inline"><mml:mo>↑</mml:mo></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col4"><inline-formula><mml:math id="M248" display="inline"><mml:mo>↑</mml:mo></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col5">The soil's high initial saturation combined with increasing spring precipitation inputs exceeds the remaining storage capacity, temporarily increasing subsurface runoff.</oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1">Late summer/fall</oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M249" display="inline"><mml:mo>↑</mml:mo></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3"><inline-formula><mml:math id="M250" display="inline"><mml:mo>↑</mml:mo></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col4">No difference</oasis:entry>
         <oasis:entry colname="col5">A transitional period as soil moisture recovers from the summer peak; no difference in subsurface runoff.</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Late fall/winter</oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M251" display="inline"><mml:mrow><mml:mo>↑</mml:mo><mml:mo>↑</mml:mo></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3">No difference</oasis:entry>
         <oasis:entry colname="col4"><inline-formula><mml:math id="M252" display="inline"><mml:mo>↓</mml:mo></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col5">Low ET allows the enhanced retention capacity to maximize SM recovery, reducing subsurface runoff under SOC scenarios.</oasis:entry>
       </oasis:row>
     </oasis:tbody>
   </oasis:tgroup></oasis:table></table-wrap>

      <p id="d2e4340">Table <xref ref-type="table" rid="T4"/> shows the relative changes for ET, total soil water content, and subsurface runoff across all cells in each subcatchment and the entire catchment over all scenarios. While overall changes are small, the signal of increased SOC is clearly visible at the catchment scale, largely due to the dominance of agricultural land. Among subcatchments, Petit Glâne and Arbogne show the largest relative changes for all variables, whereas Flon and Broye subcatchment exhibit smaller responses. Subsurface runoff at the catchment scale corresponds to the river discharge; thus, the changes in this variable reflect the impact of SOC increases on overall catchment discharge.</p>

<table-wrap id="T4" specific-use="star"><label>Table 4</label><caption><p id="d2e4349">Relative changes (over all scenarios) for key variables per subcatchment and overall domain. Changes in SWC are shown here integrated over the entire soil column (120 cm); although the overall change appears modest, differences are larger in the topsoil and smaller at depth (see Figs. <xref ref-type="fig" rid="F7"/> and   <xref ref-type="fig" rid="F12"/>).</p></caption><oasis:table frame="topbot"><oasis:tgroup cols="4">
     <oasis:colspec colnum="1" colname="col1" align="left"/>
     <oasis:colspec colnum="2" colname="col2" align="center"/>
     <oasis:colspec colnum="3" colname="col3" align="center"/>
     <oasis:colspec colnum="4" colname="col4" align="center"/>
     <oasis:thead>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1">Catchment</oasis:entry>
         <oasis:entry colname="col2">ET</oasis:entry>
         <oasis:entry colname="col3">SWC</oasis:entry>
         <oasis:entry colname="col4">Total subsurface runoff</oasis:entry>
       </oasis:row>
     </oasis:thead>
     <oasis:tbody>
       <oasis:row>
         <oasis:entry colname="col1">Arbogne</oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M253" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>0.18 % to <inline-formula><mml:math id="M254" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>0.44 %</oasis:entry>
         <oasis:entry colname="col3"><inline-formula><mml:math id="M255" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>1.43 to <inline-formula><mml:math id="M256" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>3.93 %</oasis:entry>
         <oasis:entry colname="col4"><inline-formula><mml:math id="M257" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.55 to <inline-formula><mml:math id="M258" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>1.40 %</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Broye</oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M259" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>0.13 % to <inline-formula><mml:math id="M260" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>0.32 %</oasis:entry>
         <oasis:entry colname="col3"><inline-formula><mml:math id="M261" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>1.25 to <inline-formula><mml:math id="M262" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>3.56 %</oasis:entry>
         <oasis:entry colname="col4"><inline-formula><mml:math id="M263" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.21 to <inline-formula><mml:math id="M264" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.54 %</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Flon</oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M265" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>0.09 % to <inline-formula><mml:math id="M266" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>0.24 %</oasis:entry>
         <oasis:entry colname="col3"><inline-formula><mml:math id="M267" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>1.34 to <inline-formula><mml:math id="M268" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>3.92 %</oasis:entry>
         <oasis:entry colname="col4"><inline-formula><mml:math id="M269" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.11 to <inline-formula><mml:math id="M270" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.31 %</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Petit Glâne</oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M271" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>0.27 % to <inline-formula><mml:math id="M272" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>0.67 %</oasis:entry>
         <oasis:entry colname="col3"><inline-formula><mml:math id="M273" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>1.72 to <inline-formula><mml:math id="M274" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>4.72 %</oasis:entry>
         <oasis:entry colname="col4"><inline-formula><mml:math id="M275" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.69 to <inline-formula><mml:math id="M276" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>1.74 %</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Overall</oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M277" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>0.15 % to <inline-formula><mml:math id="M278" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>0.38 %</oasis:entry>
         <oasis:entry colname="col3"><inline-formula><mml:math id="M279" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>1.34 to <inline-formula><mml:math id="M280" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>3.75 %</oasis:entry>
         <oasis:entry colname="col4"><inline-formula><mml:math id="M281" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.27 to <inline-formula><mml:math id="M282" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.70 %</oasis:entry>
       </oasis:row>
     </oasis:tbody>
   </oasis:tgroup></oasis:table></table-wrap>

</sec>
<sec id="Ch1.S3.SS4">
  <label>3.4</label><title>Impact of increasing SOC on discharge and hydrological extremes</title>
      <p id="d2e4669">Beyond grid-scale subsurface, mHM also simulates routed discharge at the locations of gauging stations. The overall effect of increased SOC on discharge is small. Because hydrographs from all scenarios almost entirely overlap, they are shown only in the Sect. S6. Relative differences between the base and SOC scenarios are moderate (positive values indicate higher discharge under SOC; Fig. <xref ref-type="fig" rid="F9"/>), which is consistent with the small changes in subsurface runoff at the grid-cell scale (Sect. <xref ref-type="sec" rid="Ch1.S3.SS3"/>).</p>

      <fig id="F9" specific-use="star"><label>Figure 9</label><caption><p id="d2e4678">Relative difference timeseries of SOC to base scenario. positive values = more discharge under SOC increase. Gray vertical lines <inline-formula><mml:math id="M283" display="inline"><mml:mo>=</mml:mo></mml:math></inline-formula> days where the low flow threshold for each subcatchment is reached in the base scenario, red vertical lines <inline-formula><mml:math id="M284" display="inline"><mml:mo>=</mml:mo></mml:math></inline-formula> days where <inline-formula><mml:math id="M285" display="inline"><mml:mrow><mml:msub><mml:mi>Q</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> threshold is reached.</p></caption>
          <graphic xlink:href="https://hess.copernicus.org/articles/30/2879/2026/hess-30-2879-2026-f09.png"/>

        </fig>

      <p id="d2e4712">All subcatchments display a similar seasonal pattern: the relative difference in discharge decreases mostly in fall and winter, and increases in spring and summer. However, the magnitude and direction of changes differ by subcatchment (Figs. <xref ref-type="fig" rid="F9"/>b and  <xref ref-type="fig" rid="F10"/>). Across catchments, relative discharge responses vary in both magnitude and direction. The Flon shows the strongest increases, with values reaching up to <inline-formula><mml:math id="M286" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>20 %, while the Petit Glâne exhibits the largest decreases of up to <inline-formula><mml:math id="M287" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>18 %. For most catchments, relative differences remain within <inline-formula><mml:math id="M288" display="inline"><mml:mrow><mml:mo>±</mml:mo><mml:mn mathvariant="normal">10</mml:mn></mml:mrow></mml:math></inline-formula> %. In absolute terms, the Arbogne (mean discharge (mean <inline-formula><mml:math id="M289" display="inline"><mml:mi>Q</mml:mi></mml:math></inline-formula>) 0.73 <inline-formula><mml:math id="M290" display="inline"><mml:mrow class="unit"><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mn mathvariant="normal">3</mml:mn></mml:msup><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">s</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula>) shows increases of up to 0.08 <inline-formula><mml:math id="M291" display="inline"><mml:mrow class="unit"><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mn mathvariant="normal">3</mml:mn></mml:msup><mml:mspace width="0.125em" linebreak="nobreak"/><mml:msup><mml:mi mathvariant="normal">s</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula> and decreases between 0.10 and 0.30 <inline-formula><mml:math id="M292" display="inline"><mml:mrow class="unit"><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mn mathvariant="normal">3</mml:mn></mml:msup><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">s</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula>. In the Broye subcatchment (mean <inline-formula><mml:math id="M293" display="inline"><mml:mi>Q</mml:mi></mml:math></inline-formula> 7.89 <inline-formula><mml:math id="M294" display="inline"><mml:mrow class="unit"><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mn mathvariant="normal">3</mml:mn></mml:msup><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">s</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula>), discharge can increase by up to 0.8 <inline-formula><mml:math id="M295" display="inline"><mml:mrow class="unit"><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mn mathvariant="normal">3</mml:mn></mml:msup><mml:mspace width="0.125em" linebreak="nobreak"/><mml:msup><mml:mi mathvariant="normal">s</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula> and decrease by <inline-formula><mml:math id="M296" display="inline"><mml:mrow><mml:mn mathvariant="normal">1</mml:mn><mml:mo>-</mml:mo><mml:mn mathvariant="normal">3.5</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M297" display="inline"><mml:mrow class="unit"><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mn mathvariant="normal">3</mml:mn></mml:msup><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">s</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula>. For the Flon (mean <inline-formula><mml:math id="M298" display="inline"><mml:mi>Q</mml:mi></mml:math></inline-formula> 0.34 <inline-formula><mml:math id="M299" display="inline"><mml:mrow class="unit"><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mn mathvariant="normal">3</mml:mn></mml:msup><mml:mspace width="0.125em" linebreak="nobreak"/><mml:msup><mml:mi mathvariant="normal">s</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula>), increases reach up to 0.05 <inline-formula><mml:math id="M300" display="inline"><mml:mrow class="unit"><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mn mathvariant="normal">3</mml:mn></mml:msup><mml:mspace width="0.125em" linebreak="nobreak"/><mml:msup><mml:mi mathvariant="normal">s</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula>, while decreases range from 0.05 to 0.25 <inline-formula><mml:math id="M301" display="inline"><mml:mrow class="unit"><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mn mathvariant="normal">3</mml:mn></mml:msup><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">s</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula>. In the Petit Glâne (mean <inline-formula><mml:math id="M302" display="inline"><mml:mi>Q</mml:mi></mml:math></inline-formula> 0.94 <inline-formula><mml:math id="M303" display="inline"><mml:mrow class="unit"><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mn mathvariant="normal">3</mml:mn></mml:msup><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">s</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula>), discharge increases are around 0.1 <inline-formula><mml:math id="M304" display="inline"><mml:mrow class="unit"><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mn mathvariant="normal">3</mml:mn></mml:msup><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">s</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula>, and decreases range from 0.3 to 0.8 <inline-formula><mml:math id="M305" display="inline"><mml:mrow class="unit"><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mn mathvariant="normal">3</mml:mn></mml:msup><mml:mspace width="0.125em" linebreak="nobreak"/><mml:msup><mml:mi mathvariant="normal">s</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula> (see Appendix <xref ref-type="sec" rid="App1.Ch1.S3.SS2"/>).</p>

      <fig id="F10" specific-use="star"><label>Figure 10</label><caption><p id="d2e5032">Spatial patterns of precipitation, temperature and discharge for one intermediate SOC scenario (MedC_lowC). Difference in discharge = SOC-scenario – base scenario. More details and monthly maps in Appendix <xref ref-type="sec" rid="App1.Ch1.S3.SS5"/>.</p></caption>
          <graphic xlink:href="https://hess.copernicus.org/articles/30/2879/2026/hess-30-2879-2026-f10.png"/>

        </fig>

      <p id="d2e5043">The Arbogne resembles the Petit Glâne, and the response of the Broye subcatchment is more attenuated. The share of pervious area per catchment (where SOC is increased) is comparable between the subcatchments: The Broye subcatchment and Arbogne have 61 % and 62 %, Flon and Petit Glâne slightly higher shares with 72 % and 74 % (Fig. <xref ref-type="fig" rid="F3"/>). Thus, the described differences between stations rather arise from climatic variations than differences in land use.</p>
      <p id="d2e5048">The Flon chatchment has a higher average elevation, with lower temperatures, more precipitation and therefore increased discharge (Fig. <xref ref-type="fig" rid="F10"/>). The Petit Glâne and Arbogne lie lower and receive less precipitation and show therefore also less discharge. The Broye subcatchment spans a wider elevational and climatic gradient, thus slightly averaging out the effects.</p>
      <p id="d2e5053">Peak flows are, in general, reduced under the SOC scenarios, although the effect is small (Fig. <xref ref-type="fig" rid="F9"/>). Floods with a 2-year return period occurred in winter 2017/2018 and summer 2021 (<inline-formula><mml:math id="M306" display="inline"><mml:mrow><mml:msub><mml:mi>Q</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> events, red vertical lines in Fig. <xref ref-type="fig" rid="F9"/>). Discharge during these events is slightly decreased under the SOC scenarios in 2017/2018, but the impact in 2021 is negligible. For instance, the peak flow in the Broye subcatchment in 2018 reached <inline-formula><mml:math id="M307" display="inline"><mml:mrow><mml:mn mathvariant="normal">77.17</mml:mn><mml:mspace width="0.125em" linebreak="nobreak"/><mml:mrow class="unit"><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mn mathvariant="normal">3</mml:mn></mml:msup><mml:mspace width="0.125em" linebreak="nobreak"/><mml:msup><mml:mi mathvariant="normal">s</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:mrow></mml:math></inline-formula> and was reduced by <inline-formula><mml:math id="M308" display="inline"><mml:mn mathvariant="normal">0.2</mml:mn></mml:math></inline-formula>–<inline-formula><mml:math id="M309" display="inline"><mml:mrow><mml:mn mathvariant="normal">0.5</mml:mn><mml:mspace linebreak="nobreak" width="0.125em"/><mml:mrow class="unit"><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mn mathvariant="normal">3</mml:mn></mml:msup><mml:mspace width="0.125em" linebreak="nobreak"/><mml:msup><mml:mi mathvariant="normal">s</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:mrow></mml:math></inline-formula> under the SOC scenarios, while in 2021 a peak of <inline-formula><mml:math id="M310" display="inline"><mml:mrow><mml:mn mathvariant="normal">70.29</mml:mn><mml:mspace linebreak="nobreak" width="0.125em"/><mml:mrow class="unit"><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mn mathvariant="normal">3</mml:mn></mml:msup><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">s</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:mrow></mml:math></inline-formula> was reduced by <inline-formula><mml:math id="M311" display="inline"><mml:mn mathvariant="normal">0.3</mml:mn></mml:math></inline-formula>–<inline-formula><mml:math id="M312" display="inline"><mml:mrow><mml:mn mathvariant="normal">1</mml:mn><mml:mspace linebreak="nobreak" width="0.125em"/><mml:mrow class="unit"><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mn mathvariant="normal">3</mml:mn></mml:msup><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">s</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:mrow></mml:math></inline-formula> (see Appendix <xref ref-type="sec" rid="App1.Ch1.S3.SS6"/>).</p>
      <p id="d2e5184">A relevant indicator for low flows is the <inline-formula><mml:math id="M313" display="inline"><mml:mrow><mml:msub><mml:mi>Q</mml:mi><mml:mn mathvariant="normal">347</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> threshold, an indicator that cantonal authorities use to determine bans on irrigation water withdrawal from rivers to fulfill the minimum environmental flow requirements. In Fig. <xref ref-type="fig" rid="F9"/>, days where the observed discharge fell below this threshold are marked in gray as low-flow days. Although the influence is minor, discharge is slightly increased under the SOC scenarios before and sometimes during observed low-flow periods. This leads to fewer days falling below the <inline-formula><mml:math id="M314" display="inline"><mml:mrow><mml:msub><mml:mi>Q</mml:mi><mml:mn mathvariant="normal">347</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> – typically <inline-formula><mml:math id="M315" display="inline"><mml:mn mathvariant="normal">1</mml:mn></mml:math></inline-formula>–<inline-formula><mml:math id="M316" display="inline"><mml:mn mathvariant="normal">6</mml:mn></mml:math></inline-formula> d depending on scenario, year, and subcatchment (in the Broye subcatchment, for example, <inline-formula><mml:math id="M317" display="inline"><mml:mn mathvariant="normal">1</mml:mn></mml:math></inline-formula>–<inline-formula><mml:math id="M318" display="inline"><mml:mn mathvariant="normal">4</mml:mn></mml:math></inline-formula> d less). However, in the Arbogne in 2016 and 2019, as well as in the Petit Glâne in 2019, low-flow periods coincided with reduced discharge under the SOC scenarios, resulting in more low-flow days (a surplus of <inline-formula><mml:math id="M319" display="inline"><mml:mn mathvariant="normal">1</mml:mn></mml:math></inline-formula>–<inline-formula><mml:math id="M320" display="inline"><mml:mn mathvariant="normal">14</mml:mn></mml:math></inline-formula> d in the Arbogne and up to <inline-formula><mml:math id="M321" display="inline"><mml:mn mathvariant="normal">5</mml:mn></mml:math></inline-formula> d in the Petit Glâne, Fig. <xref ref-type="fig" rid="F11"/>).</p>

      <fig id="F11"><label>Figure 11</label><caption><p id="d2e5266">Timeseries of relative difference between base and SOC scenarios in the annual number of days with discharge below <inline-formula><mml:math id="M322" display="inline"><mml:mrow><mml:msub><mml:mi>Q</mml:mi><mml:mn mathvariant="normal">347</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> (low flow threshold).</p></caption>
          <graphic xlink:href="https://hess.copernicus.org/articles/30/2879/2026/hess-30-2879-2026-f11.png"/>

        </fig>


</sec>
<sec id="Ch1.S3.SS5">
  <label>3.5</label><title>Scenario sensitivity</title>
      <p id="d2e5297">Since the overall small impact of SOC increase on ET was first surprising, we wanted to investigate the responses of the individual soil layers. Here we found, that although the soil water content in the first two layers was consistently higher under the SOC scenarios, ET from soil layers 1 is reduced, while it is increased from soil layer 2 and 3, leading to an overall small net increase. The reason for this is explained and discussed in Sect. <xref ref-type="sec" rid="Ch1.S4.SS2.SSS3"/>.</p>
      <p id="d2e5302">The SOC scenarios represent possible outcomes of combinations of management adaptations. Their impact on the model output fluxes ET and total grid-scale runoff increases almost linearly with increasing SOC content, as visible by comparing scenarios LowC_vLowC, MedC_LowC and vHighC_MedC in Fig. <xref ref-type="fig" rid="F12"/>.</p>

      <fig id="F12"><label>Figure 12</label><caption><p id="d2e5309">Magnitude of change in selected states and fluxes for SOC scenarios relative to the base scenario.</p></caption>
          <graphic xlink:href="https://hess.copernicus.org/articles/30/2879/2026/hess-30-2879-2026-f12.png"/>

        </fig>

      <p id="d2e5319">In total, the largest SOC additions occur in scenario vHighC_MedC (<inline-formula><mml:math id="M323" display="inline"><mml:mo lspace="0mm">+</mml:mo></mml:math></inline-formula>1.5 % in the first layer and <inline-formula><mml:math id="M324" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>0.6 % in the second layer), whereas in scenario MedC_MedC, SOC is added evenly across both layers (<inline-formula><mml:math id="M325" display="inline"><mml:mo lspace="0mm">+</mml:mo></mml:math></inline-formula>1 % in each). Interestingly, the effects on soil moisture are often largest under MedC_MedC, despite its slightly lower total SOC increase (Fig. <xref ref-type="fig" rid="F7"/>). This suggests that increasing SOC in the subsoil can be particularly beneficial, as it slows soil moisture depletion during late summer and fall. Differences between vHighC_MedC and MedC_MedC are generally minor: in most catchments, the two scenarios produce nearly identical reductions in low-flow days compared to the base scenario. However, in the Arbogne in 2018, low-flow days are reduced by one day only under MedC_MedC, and in the Flon in 2022, the reduction under scenario MedC_MedC is two days – one day more than under vHighC_MedC.</p>
      <p id="d2e5345">For the catchment-wide, annual effect, the distribution of SOC in the two soil layers does not make a difference. Only at the seasonal scale, a distribution into deeper layers might lead to a delay of drought-induced transpiration reduction (as was observed in <xref ref-type="bibr" rid="bib1.bibx121 bib1.bibx49" id="altparen.117"/>).</p>
</sec>
</sec>
<sec id="Ch1.S4">
  <label>4</label><title>Discussion</title>
<sec id="Ch1.S4.SS1">
  <label>4.1</label><title>Applicability of the study framework</title>
<sec id="Ch1.S4.SS1.SSS1">
  <label>4.1.1</label><title>Model performance</title>
      <p id="d2e5375">In this study, we used a fixed parameter set that was calibrated to perform well across all four subcatchments, effectively representing observed discharge dynamics. Model or parameter uncertainty was not systematically explored, but testing six alternative parameter sets showed that the direction of simulated changes is robust, while the magnitude varies (Appendix <xref ref-type="sec" rid="App1.Ch1.S4"/>).</p>
      <p id="d2e5380">For streamflow, the calibrated mHM model performs very well for the Broye subcatchment (KGE <inline-formula><mml:math id="M326" display="inline"><mml:mo>=</mml:mo></mml:math></inline-formula> 0.91, NSE <inline-formula><mml:math id="M327" display="inline"><mml:mo>=</mml:mo></mml:math></inline-formula> 0.86), outperforming previous applications of conceptual models (SWAT, <xref ref-type="bibr" rid="bib1.bibx135" id="altparen.118"/>; PREVAH, <xref ref-type="bibr" rid="bib1.bibx81" id="altparen.119"/>) and a physics-based model (Alpine3D, <xref ref-type="bibr" rid="bib1.bibx68" id="altparen.120"/>). Despite relatively short calibration and evaluation periods (four and three years), these performance values are high (see Sect. S1 for a more detailed comparison), underlining the model's ability to reproduce observed discharge dynamics. Seasonal low-flow regimes are fairly well reproduced for the Broye subcatchment and Petit Glâne, while the frequency of low flows is underestimated for the Flon and overestimated for the Arbogne. The differences in performance can be traced to biases in the precipitation input fields. Such biases were already reported in earlier studies using the same precipitation data product <xref ref-type="bibr" rid="bib1.bibx15 bib1.bibx81" id="paren.121"/>. While our adjustments to the precipitation input substantially reduced these biases, they were not fully eliminated. A more systematic bias correction would be required. Still, this adaptation was essential to reliably simulate soil moisture dynamics. Here, we were able to reproduce observed soil moisture time series with relatively good performance (Appendix <xref ref-type="sec" rid="App1.Ch1.S3.SS1"/>).</p>
</sec>
<sec id="Ch1.S4.SS1.SSS2">
  <label>4.1.2</label><title>Plausibility of represented changes in soil hydraulic properties</title>
      <p id="d2e5420">The plausibility of our simulation results depends on how reliably SOC-driven changes in soil hydraulic properties are represented, which can only be discussed against literature reported estimates.</p>
      <p id="d2e5423">The non-linear pedotransfer function (PTF) used to calculate <inline-formula><mml:math id="M328" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi mathvariant="normal">b</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> from SOC captures the stronger sensitivity of <inline-formula><mml:math id="M329" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">FC</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> and <inline-formula><mml:math id="M330" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">Sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> at low initial SOC, consistent with large soil database analyses <xref ref-type="bibr" rid="bib1.bibx52 bib1.bibx93 bib1.bibx77" id="paren.122"/>. Simulated reductions in <inline-formula><mml:math id="M331" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi mathvariant="normal">b</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> (Fig. <xref ref-type="fig" rid="F6"/>) align with reported ranges (Table <xref ref-type="table" rid="T1"/>). In our study, <inline-formula><mml:math id="M332" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">FC</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> and <inline-formula><mml:math id="M333" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">Sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> increase with SOC, while <inline-formula><mml:math id="M334" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">PWP</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> increases less, raising plant available water capacity (PAWC), in line with previous findings <xref ref-type="bibr" rid="bib1.bibx92 bib1.bibx93 bib1.bibx70 bib1.bibx66 bib1.bibx1" id="paren.123"/>. Exact incremental changes along the retention curve remain however uncertain and are soil-specific <xref ref-type="bibr" rid="bib1.bibx66" id="paren.124"/>. Reported PAWC increases vary widely: 1.5 %–7 % for <inline-formula><mml:math id="M335" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>0.6 % SOC <xref ref-type="bibr" rid="bib1.bibx84 bib1.bibx70" id="paren.125"/>, 1.16 % for <inline-formula><mml:math id="M336" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>1 % SOC <xref ref-type="bibr" rid="bib1.bibx77" id="paren.126"/>, up to 50 % for <inline-formula><mml:math id="M337" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>1.5 % SOC <xref ref-type="bibr" rid="bib1.bibx70" id="paren.127"/>, and 4 %–45 % for management-related SOC increases by 7 %–220 % <xref ref-type="bibr" rid="bib1.bibx46 bib1.bibx45 bib1.bibx10" id="paren.128"/>. Our simulated average increase of 9.3 % for <inline-formula><mml:math id="M338" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>1 % SOC (<inline-formula><mml:math id="M339" display="inline"><mml:mo lspace="0mm">+</mml:mo></mml:math></inline-formula>1 % (mass) increase <inline-formula><mml:math id="M340" display="inline"><mml:mo>≈</mml:mo></mml:math></inline-formula> 35 %–60 % relative increase) lies within these ranges but toward the upper end. Changes in <inline-formula><mml:math id="M341" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">Sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> or <inline-formula><mml:math id="M342" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">FC</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> are rarely quantified, but <xref ref-type="bibr" rid="bib1.bibx104" id="text.129"/> reported <inline-formula><mml:math id="M343" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>8 %–10 % in a silt loam, comparable to our <inline-formula><mml:math id="M344" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>6.2 % average. Changes in <inline-formula><mml:math id="M345" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">PWP</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> are unfortunately rarely reported, meaning that we cannot explicitly assess the plausibility of our simulated average increase in <inline-formula><mml:math id="M346" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">PWP</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> by 2.1 %. The SOC effect on water retention is texture dependent, with greater PAWC increases in coarser soils <xref ref-type="bibr" rid="bib1.bibx70 bib1.bibx66" id="paren.130"/> and at low initial SOC <xref ref-type="bibr" rid="bib1.bibx93" id="paren.131"/>, patterns consistent with our results (see Sect. S7).</p>
      <p id="d2e5642">The simulated change of <inline-formula><mml:math id="M347" display="inline"><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> aligns with observations from <xref ref-type="bibr" rid="bib1.bibx46 bib1.bibx45 bib1.bibx10" id="text.132"/> for similar soils, is higher than those reported by <xref ref-type="bibr" rid="bib1.bibx93 bib1.bibx125" id="text.133"/>, but comparable to <xref ref-type="bibr" rid="bib1.bibx13 bib1.bibx60" id="text.134"/>, who also used PTF estimates rather than observations, which carry their own assumptions and uncertainties. Given high variability in pedo-climatic conditions, management practices, PTF selection, and soil texture, the plausibility of simulated hydraulic changes can only be assessed generally: overall trends and magnitudes are plausible, but uncertainty remains, especially for <inline-formula><mml:math id="M348" display="inline"><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> and <inline-formula><mml:math id="M349" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">PWP</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>. A potential way forward would be to incorporate additional soil probe measurements and observationally derived hydraulic parameters, such as <inline-formula><mml:math id="M350" display="inline"><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, to further constrain and validate the model.</p>
</sec>
</sec>
<sec id="Ch1.S4.SS2">
  <label>4.2</label><title>Simulated impact of management adaptations</title>
<sec id="Ch1.S4.SS2.SSS1">
  <label>4.2.1</label><title>SOC enhances ET and reduces subsurface runoff at the grid-scale</title>
      <p id="d2e5715">The SOC scenarios considered here assume uniform application of SOC across all pervious landcover cells (arable land). Despite this simplification, their impact on simulated hydrological processes is small. This can be attributed to the moderate changes in hydraulic properties following SOC increases, which fall within observed ranges and can therefore be considered plausible (Sect. <xref ref-type="sec" rid="Ch1.S4.SS1.SSS2"/>). Across scenarios, soil water content (<inline-formula><mml:math id="M351" display="inline"><mml:mi mathvariant="italic">θ</mml:mi></mml:math></inline-formula>) in the first two layers increases by 8.8 % to 3.2 % (Fig. <xref ref-type="fig" rid="F7"/>). This increased retention capacity slightly reducing percolation and temporarily enabling higher ET during summer when evaporative demand peaks and when ET would otherwise be water-limited. Over 2016–2022, this results in a net ET increase of <inline-formula><mml:math id="M352" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>0.16 %–0.4 % (8–18 mm depending on scenario, Fig. <xref ref-type="fig" rid="F7"/>). When aggregated over the entire catchment, the relative changes in soil moisture and ET are slightly smaller, but the signal remains clearly visible due to the high fraction of arable land across all subcatchments.</p>
      <p id="d2e5738">Direct comparison with experiments is difficult since ET is rarely measured in field experiments; PAWC is often used as a proxy due to its influence on transpiration and yield <xref ref-type="bibr" rid="bib1.bibx38" id="paren.135"/>. Across European sites, although overall impacts on water retention were limited, SOC increases modestly raised PAWC and slightly delayed plant drought stress <xref ref-type="bibr" rid="bib1.bibx108" id="paren.136"/>. This implies a modest rise in transpiration, but it was not measured directly. A lysimeter experiment showed that biochar application to sandy soil reduced bulk density, increased porosity, and ultimately enhanced ET <xref ref-type="bibr" rid="bib1.bibx42" id="paren.137"/>.</p>
      <p id="d2e5750">Plot-scale simulations with Richards-equation-based models show that <inline-formula><mml:math id="M353" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>1 % (mass) SOC can increase transpiration by up to 9 %, while soil evaporation may decrease due to higher crop cover <xref ref-type="bibr" rid="bib1.bibx49" id="paren.138"/>. <xref ref-type="bibr" rid="bib1.bibx121" id="text.139"/> reported similar increases (<inline-formula><mml:math id="M354" display="inline"><mml:mo lspace="0mm">+</mml:mo></mml:math></inline-formula>15 mm yr<sup>−1</sup> for <inline-formula><mml:math id="M356" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>1 % SOC down to 65 cm). Using Hydrus-1D <xref ref-type="bibr" rid="bib1.bibx107" id="paren.140"/>, <xref ref-type="bibr" rid="bib1.bibx38" id="text.141"/> found that evaporation always increased, whereas transpiration rose only when SOC increased below 30 cm depth, with stronger effects in finer soils and drought years. Deep drainage and recharge consistently declined, in line with our simulated percolation and recharge decreases.</p>
      <p id="d2e5799">Unlike these plot-scale studies, mHM represents ET as a single bulk flux after canopy interception, without separating evaporation and transpiration or explicitly limiting root water uptake. Consequently, direct comparison with plot-scale studies is limited. We argue, the simulated ET increase in this study is likely dominated by transpiration, which the increase in PAWC suggests (Fig. <xref ref-type="fig" rid="F6"/>). During high summer evaporative demand, the increased PAWC and <inline-formula><mml:math id="M357" display="inline"><mml:mi mathvariant="italic">θ</mml:mi></mml:math></inline-formula> translate directly into higher ET, slightly reducing subsurface runoff (<inline-formula><mml:math id="M358" display="inline"><mml:mo lspace="0mm">-</mml:mo></mml:math></inline-formula>0.3 % to <inline-formula><mml:math id="M359" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.78 %, 10–22 mm over 7 years).</p>
      <p id="d2e5826">SOC impacts on ET strongly depend on management: cover crops, mulching, or residue retention can suppress soil evaporation <xref ref-type="bibr" rid="bib1.bibx1" id="paren.142"/>. The effect of topsoil changes is two-fold: soil cover (like mulching) can (i) modify the re-evaporation of (soil-)intercepted water and (ii) affect soil evaporation of infiltrated water <xref ref-type="bibr" rid="bib1.bibx91" id="paren.143"/>. While the first could, in principle, be captured bymodified interception parameterizations, the second remains largely absent from current catchment-scale models and represents an direction for future work – explicitly distinguishing soil evaporation and transpiration across temporal and vertical scales.</p>
      <p id="d2e5835">In summary, modest SOC increases slightly enhance summer ET, likely via transpiration, and marginally reduce subsurface runoff. These effects align with field-scale findings and modeling results, particularly where SOC increases extend deeper into the soil, underscoring the importance of considering both depth and method of SOC application in agricultural practice. The simulated reduction of deeper drainage, and thus recharge, in our and other modeling studies highlights a potential trade-off between enhancing SOC for agricultural benefits and sustaining hydrologic processes critical for water management, especially under changing climate conditions.</p>
      <p id="d2e5838">Although it is beyond the scope of this study, we acknowledge that increasing SOC also affects soil biogeochemical cycles, particularly when nutrient balances change. For example, in poorly-drained soils, increasing SOC without adjusting nitrogen inputs can enhance denitrification and lead to elevated emissions of N<sub>2</sub>O, a potent greenhouse gas, thus representing a trade-off worth noting <xref ref-type="bibr" rid="bib1.bibx55" id="paren.144"/>.</p>
</sec>
<sec id="Ch1.S4.SS2.SSS2">
  <label>4.2.2</label><title>Catchment-scale implications of SOC-induced changes in discharge</title>
      <p id="d2e5861">SOC-related impacts on discharge are seasonally dependent. In spring, increased rainfall combined with high soil moisture (<inline-formula><mml:math id="M361" display="inline"><mml:mi mathvariant="italic">θ</mml:mi></mml:math></inline-formula>) levels from winter, pushing more water into percolation and discharge just before the low-flow period. During the low-flow period itself, higher ET reduces percolation, so discharge increases are limited (1 %–5 %) or may even turn into decreases. These modest discharge increases can reduce days below the <inline-formula><mml:math id="M362" display="inline"><mml:mrow><mml:msub><mml:mi>Q</mml:mi><mml:mn mathvariant="normal">347</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> threshold by 1–6 compared to the base scenario, potentially easing irrigation constraints <xref ref-type="bibr" rid="bib1.bibx49" id="paren.145"/>. This holds for the Flon and Broye subcatchment, but in the Arbogne (2016, 2019) and Petit Glâne (2017, 2019), low-flow days mostly increase. These two subcatchments are the lowest, thus warmer and also drier than the others. When low subsurface runoff coincides with high ET, the SOC scenarios further enhance water retention and ET (as visible in Table <xref ref-type="table" rid="T4"/>), which can exacerbate discharge reductions and increase low-flow days, potentially increasing the likelihood of irrigation constraints.</p>
      <p id="d2e5887">Seasonal discharge dynamics are best captured by the model in the Broye subcatchment; smaller subcatchments show biased or mis-timed low flows, likely reflecting input data limitations. Changes in low-flow days, derived from the 95th percentile of discharge, are sensitive to model optimization and thus less robust. While overall discharge fits are nearly identical across six optimization runs, variation at the distribution tails is observed and expected (Appendix <xref ref-type="sec" rid="App1.Ch1.S4"/>). The selected optimization run reflects the overall pattern observed over several optimization runs: reductions in low-flow days are consistent in the Broye subcatchment and Flon, whereas the pattern for the Arbogne and Petit Glâne are more variable. However, the number of low flow days most often increases in 2016, 2017 and 2019. This suggests that SOC increases likely reduce the number of low-flow days in larger or cooler, wetter catchments. However, the impacts in smaller, warmer, drier catchments are highly variable. Often, they even increase the number of low flows. Small catchments, with typically low storage and fast hydrological response, are highly sensitive to minor changes in precipitation and temperature <xref ref-type="bibr" rid="bib1.bibx118" id="paren.146"/>. Thus, even modest SOC-induced reductions in discharge can push flows below ecological or minimum thresholds, making these trade-offs especially relevant in smaller catchments under future climate changes.</p>
      <p id="d2e5895">Observed discharge in the Broye subcatchment peaks between December and March, but SOC-related reductions occur mainly in late autumn and winter, not necessarily the most critical periods (Fig. <xref ref-type="fig" rid="F9"/>). <inline-formula><mml:math id="M363" display="inline"><mml:mrow><mml:msub><mml:mi>Q</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> events in winter 2017/2018 and summer 2021 show only moderate reductions (Appendix <xref ref-type="sec" rid="App1.Ch1.S3.SS6"/>). Daily simulation resolution prevents precise quantification of peak reductions.</p>
      <p id="d2e5913">Evidence on agricultural management effects on peak and low flows is limited, with most studies focusing on major land-use changes or structural interventions. As a result, direct validation of our findings is challenging, and comparisons must be made cautiously. In a modeling experiment on land use changes, <xref ref-type="bibr" rid="bib1.bibx80" id="text.147"/> found that high <inline-formula><mml:math id="M364" display="inline"><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> reduces mean discharge and flood peaks, consistent with our findings, although we only found a very limited effect of increased <inline-formula><mml:math id="M365" display="inline"><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> on <inline-formula><mml:math id="M366" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">FC</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>. <xref ref-type="bibr" rid="bib1.bibx2" id="text.148"/> simulated the impact of cover crops and reduced tillage and found a moderate reduction in high-frequency flood peaks, also in line with our results. Similarly, <xref ref-type="bibr" rid="bib1.bibx34" id="text.149"/> found that strong soil compaction at the plot scale (represented by <inline-formula><mml:math id="M367" display="inline"><mml:mi mathvariant="normal">−</mml:mi></mml:math></inline-formula>95 % <inline-formula><mml:math id="M368" display="inline"><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>) can increase discharge peaks by 50 %–80 % at the catchment scale, as shown in their study based on the Tethys-Chloris model.</p>
      <p id="d2e5978">The aim of this study was to explore sensitivities of catchment-scale hydrological processes to increases in SOC. Thereby, our scenario assumptions are aligned with targets set by recent policy agendas promoting large-scale SOC increases in the region such as the Climate Adaptation Plan 2020 by the Canton of Vaud <xref ref-type="bibr" rid="bib1.bibx17" id="paren.150"/> and the national Climate Strategy for Agriculture and Food 2050 <xref ref-type="bibr" rid="bib1.bibx11" id="paren.151"/>. We acknowledge that the uniform SOC increase is a simplification, and likely overestimates the sequestration potential at catchment scale, particularly for the Flon, where meadows and pastures dominate, which are assumed to have smaller potentials for SOC increases than arable soils. However, evidence exists that also permanent grasslands and pastures sites hold potentials for SOC increases through management adaptation <xref ref-type="bibr" rid="bib1.bibx87 bib1.bibx43 bib1.bibx57 bib1.bibx128" id="paren.152"/>. Overall, it is known that the limits to potentials for SOC increases depend on pedoclimatic and management drivers. Their quantification, however, remains challenging <xref ref-type="bibr" rid="bib1.bibx7" id="paren.153"/>.</p>
      <p id="d2e5993">Even under these assumptions, the catchment-scale effects are very modest, which is in line with <xref ref-type="bibr" rid="bib1.bibx34" id="text.154"/>, suggesting detectable impacts of management require either strong interventions or long observation periods.</p>
      <p id="d2e5999">It should be noted that conventional agriculture can lead to SOC losses, if more carbon is removed from the soil than is returned <xref ref-type="bibr" rid="bib1.bibx65 bib1.bibx101 bib1.bibx58" id="paren.155"/>. These losses are expected to accelerate under climate change, because higher temperatures enhance SOC mineralization <xref ref-type="bibr" rid="bib1.bibx25 bib1.bibx129 bib1.bibx41" id="paren.156"/>. Exploring scenarios with decreasing SOC could provide additional insights in future studies. Herein, we focus on an optimistic SOC increase scenario to illustrate potential upper-bound effects on soil water retention and catchment hydrological responses.</p>
      <p id="d2e6008">In recent years, a broader debate around nature-based solutions and soil and water conservation measures has emerged. Several international and European initiatives aim to enhance soil carbon sequestration, soil health, and water retention through nature-based and conservation practices. The 4 per mille Initiative (<uri>https://sdgs.un.org/partnerships/4-1000-initiative-and-its-implementation</uri>, last access: 10 May 2026) promotes the increase of global SOC stocks by 0.4 % yr<sup>−1</sup> to offset CO<sub>2</sub> emissions from fossil fuels <xref ref-type="bibr" rid="bib1.bibx78" id="paren.157"/>. The EU project NBsoil (<uri>https://nbsoil.eu/</uri>, last access: 10 May 2026) focuses on nature-based soil management to enhance soil ecosystem services. The EJP Soil project SoilX (<uri>https://projects.au.dk/ejpsoil/soil-research/eom4soil/into-dialogue/soilx</uri>, last access: 10 May 2026) develops strategies to improve soil carbon, soil health, and water retention. The OPTAIN project (<uri>https://www.optain.eu/</uri>, last access: 10 May 2026) promotes small water retention measures and nutrient management in agricultural catchments. While increasing SOC can enhance water retention, slightly reduce flood peaks, and decrease low-flow frequency, the catchment scale benefits remain modest even under large SOC increases. Moreover, our results indicate that in smaller, drier agricultural catchments, SOC-enhancing measures may involve trade-offs, such as reduced groundwater recharge or streamflow, reducing downstream water availability, which should be considered when designing management strategies.</p>
</sec>
<sec id="Ch1.S4.SS2.SSS3">
  <label>4.2.3</label><title>Root distribution dependency on <inline-formula><mml:math id="M371" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">FC</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> in mHM</title>
      <p id="d2e6068">As noted in Sect. <xref ref-type="sec" rid="Ch1.S3.SS5"/>, the SOC-induced increase in <inline-formula><mml:math id="M372" display="inline"><mml:mi mathvariant="italic">θ</mml:mi></mml:math></inline-formula> only led to a very small net increase in ET, since ET from the top soil layer actually decreased, which was unexpected. This response stems from the mHM adaptation by <xref ref-type="bibr" rid="bib1.bibx28" id="text.158"/>, which links root distribution to field capacity (<inline-formula><mml:math id="M373" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">FC</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>): higher <inline-formula><mml:math id="M374" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">FC</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> shifts root fractions (<inline-formula><mml:math id="M375" display="inline"><mml:mi>R</mml:mi></mml:math></inline-formula>) downward, reducing the weight of the top layer and increasing that of the lower ones. This relationship was derived from observations in the region where the scheme was developed, where sandy soils with low <inline-formula><mml:math id="M376" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">FC</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> concentrated roots near the surface, while clay-rich soils with high <inline-formula><mml:math id="M377" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">FC</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> showed deeper rooting <xref ref-type="bibr" rid="bib1.bibx28" id="paren.159"/>. Yet as noted by <xref ref-type="bibr" rid="bib1.bibx28" id="text.160"/>, such a pattern is not necessarily globally valid.</p>
      <p id="d2e6141">If we recall the formulation of the soil moisture stress function <inline-formula><mml:math id="M378" display="inline"><mml:mi>f</mml:mi></mml:math></inline-formula> (Sect. <xref ref-type="sec" rid="Ch1.S2.SS3"/>), which linearly scales PET to ET, we saw that <inline-formula><mml:math id="M379" display="inline"><mml:mi>f</mml:mi></mml:math></inline-formula> depends on the root fraction <inline-formula><mml:math id="M380" display="inline"><mml:mi>R</mml:mi></mml:math></inline-formula> and the normalized soil water content <inline-formula><mml:math id="M381" display="inline"><mml:mover accent="true"><mml:mi mathvariant="italic">θ</mml:mi><mml:mo mathvariant="normal">‾</mml:mo></mml:mover></mml:math></inline-formula> (calculated as: <inline-formula><mml:math id="M382" display="inline"><mml:mrow><mml:mover accent="true"><mml:mi mathvariant="italic">θ</mml:mi><mml:mo mathvariant="normal">‾</mml:mo></mml:mover><mml:mo>=</mml:mo><mml:mo>(</mml:mo><mml:mi mathvariant="italic">θ</mml:mi><mml:mo>-</mml:mo><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">pwp</mml:mi></mml:msub><mml:mo>)</mml:mo><mml:mo>/</mml:mo><mml:mo>(</mml:mo><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub><mml:mo>-</mml:mo><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">pwp</mml:mi></mml:msub><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula>).</p>
      <p id="d2e6223">If mean soil water content <inline-formula><mml:math id="M383" display="inline"><mml:mover accent="true"><mml:mi mathvariant="italic">θ</mml:mi><mml:mo mathvariant="normal">‾</mml:mo></mml:mover></mml:math></inline-formula> increases or decreases depends on the SOC-induced increases in <inline-formula><mml:math id="M384" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">pwp</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> and <inline-formula><mml:math id="M385" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> that are texture dependent, but also on the daily varying <inline-formula><mml:math id="M386" display="inline"><mml:mi mathvariant="italic">θ</mml:mi></mml:math></inline-formula>, which depends on incoming precipitation, so seasonality. Only if <inline-formula><mml:math id="M387" display="inline"><mml:mi mathvariant="italic">θ</mml:mi></mml:math></inline-formula> increases sufficiently, <inline-formula><mml:math id="M388" display="inline"><mml:mover accent="true"><mml:mi mathvariant="italic">θ</mml:mi><mml:mo mathvariant="normal">‾</mml:mo></mml:mover></mml:math></inline-formula> would increase and by that also <inline-formula><mml:math id="M389" display="inline"><mml:mi>f</mml:mi></mml:math></inline-formula> and hence ET. This mechanism applies in principle to both upper soil layers, but in the top layer <inline-formula><mml:math id="M390" display="inline"><mml:mi>R</mml:mi></mml:math></inline-formula> decreases as <inline-formula><mml:math id="M391" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">FC</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> increases. Consequently, even though <inline-formula><mml:math id="M392" display="inline"><mml:mover accent="true"><mml:mi mathvariant="italic">θ</mml:mi><mml:mo mathvariant="normal">‾</mml:mo></mml:mover></mml:math></inline-formula> tends to increase, the overall stress factor <inline-formula><mml:math id="M393" display="inline"><mml:mi>f</mml:mi></mml:math></inline-formula> (and thus ET) decreases in most cases. Only under very wet conditions, high soil water content <inline-formula><mml:math id="M394" display="inline"><mml:mi mathvariant="italic">θ</mml:mi></mml:math></inline-formula> may offset the reduction in root fraction and ET can still increase. In deeper layers, the opposite holds: the higher root fraction allows <inline-formula><mml:math id="M395" display="inline"><mml:mi>f</mml:mi></mml:math></inline-formula> to increase, so ET is increased when additional water infiltrates from above. Physiologically, this is unexpected, as plants usually allocate roots cost-efficiently to shallow layers where water and nutrients are accessible, though they may extend them deeper under drought <xref ref-type="bibr" rid="bib1.bibx56 bib1.bibx53 bib1.bibx40 bib1.bibx72" id="paren.161"/>. More broadly, root allocation depends on cultivar and growth stage <xref ref-type="bibr" rid="bib1.bibx114" id="paren.162"/>, and such dynamics are difficult to generalize at the catchment scale. Nevertheless, as discussed in Sect. <xref ref-type="sec" rid="Ch1.S4.SS2.SSS2"/>, the overall pattern of SOC-induced changes remains robust. Future work could test how increased <inline-formula><mml:math id="M396" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">FC</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> affects root depth allocation and evapotranspiration dynamics under local climatic and edaphic conditions in our case study region.</p>
      <p id="d2e6360">Note that in Fig. <xref ref-type="fig" rid="F12"/>, soil water content in the top layer differs slightly among the three SOC scenarios MedC_MedC, MedC_LowC and MedC_ZeroC, even though the SOC increase in this layer is identical. This results from the top-down calculation of the root fraction <inline-formula><mml:math id="M397" display="inline"><mml:mi>R</mml:mi></mml:math></inline-formula> per layer and subsequent re-normalization, which ensures that <inline-formula><mml:math id="M398" display="inline"><mml:mi>R</mml:mi></mml:math></inline-formula> sums to one across all layers. Therefore, differences in SOC in the lower soil layers can indirectly impact soil water content and ET in the top layer.</p>
</sec>
<sec id="Ch1.S4.SS2.SSS4">
  <label>4.2.4</label><title>Role of SOC increase magnitude and depth in modulating hydrological responses</title>
      <p id="d2e6387">The SOC scenarios affect ET and subsurface runoff almost linearly with increasing SOC. Seasonal differences emerge when SOC is distributed into deeper layers: Scenario MedC_MedC (<inline-formula><mml:math id="M399" display="inline"><mml:mo lspace="0mm">+</mml:mo></mml:math></inline-formula>1 % SOC in both soil layers) exhibits the highest increase in <inline-formula><mml:math id="M400" display="inline"><mml:mi mathvariant="italic">θ</mml:mi></mml:math></inline-formula> over the winter and spring, despite vHighC_MedC adding more SOC in total (Fig. <xref ref-type="fig" rid="F7"/>). Adding SOC to deeper layers delays overall soil moisture depletion and can thus reduce drought impacts, which was also concluded in the modeling studies of <xref ref-type="bibr" rid="bib1.bibx121" id="text.163"/> and <xref ref-type="bibr" rid="bib1.bibx38" id="text.164"/>. In our model simulations on catchment-annual scales, however, the vertical SOC distribution plays little role and achieving a significant increase in SOC in deeper layers is more difficult, as most (agricultural) adaptation measures would primarily lead to SOC increases near the surface <xref ref-type="bibr" rid="bib1.bibx3" id="paren.165"/>.</p>
</sec>
</sec>
<sec id="Ch1.S4.SS3">
  <label>4.3</label><title>Model suitability and structural limitations for representing SOC-induced changes</title>
      <p id="d2e6425">The mHM model is well suited for impact studies like this due to its open-source nature, active user community, and flexible structure, which allows individual adjustments, such as in the estimation of soil hydraulic properties <xref ref-type="bibr" rid="bib1.bibx71" id="paren.166"/>.</p>
      <p id="d2e6431">As with the case of any modeling scheme, some simplifications and limitations remain. First, in mHM only three land-use classes are distinguished, which may be sufficient at large scales but limits the representation of heterogeneous agricultural landscapes. In our study region, pervious land cover aggregates cropland and meadows, which can differ in management and water-use processes. Introducing additional land-use classes and distinguishing different crop functional types with varying root profiles would improve model realism. Differentiating winter crops and spring crops could be important given their distinct patterns of water uptake which may influence recharge and also low flow dynamics differently.</p>
      <p id="d2e6434">In the present framework, however, SOC effects are presented primarily through changes in soil hydraulic properties which directly control soil moisture availability and, consequently, ET. While mHM accounts for some vegetation responses (e.g., adjustments in rooting depth; Sect. <xref ref-type="sec" rid="Ch1.S4.SS2.SSS3"/>), plants are represented at a coarse level.</p>
      <p id="d2e6439">Regarding ET, mHM separates canopy interception but aggregates soil evaporation and transpiration into a single flux, as is common in many hydrological models <xref ref-type="bibr" rid="bib1.bibx95" id="paren.167"/>. While net ET is likely captured realistically, the partitioning between productive (transpiration) and unproductive (soil evaporation and interception) fluxes, as well as their temporal dynamics, may differ from reality. Finally, the root distribution, which varies with <inline-formula><mml:math id="M401" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">FC</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, is more dynamic than standard static profiles, but could be improved by incorporating dynamic crop/root growth and reassessing the negative relationship between <inline-formula><mml:math id="M402" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">FC</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> and root density under the climatic and edaphic conditions of our study region. Overall, while these limitations can affect plot scale process representation, their impact at the catchment scale is uncertain and difficult to validate, particularly given the lack of direct observations for root distribution or ET partitioning. Within these constraints, representing SOC impacts via soil hydraulic properties captures the dominant hydrological pathway relevant at the catchment scale and is therefore the focus of this study.</p>
      <p id="d2e6468">Compared to fully physics-based models such as WaSiM-ETH <xref ref-type="bibr" rid="bib1.bibx103" id="paren.168"/>, which often require extensive parameter adjustment and high computational effort, mHM offers a practical balance between spatially explicit process representation and computational efficiency <xref ref-type="bibr" rid="bib1.bibx95 bib1.bibx63 bib1.bibx96" id="paren.169"/>. Fully physics-based models are in practice never “fully” mechanistic, and for our purpose they would not provide additional advantages in representing SOC-related management effects. Their higher data and computational demands would mainly add complexity without improving the core processes relevant to this study.</p>
</sec>
</sec>
<sec id="Ch1.S5" sec-type="conclusions">
  <label>5</label><title>Conclusion</title>
      <p id="d2e6487">We explored how increasing soil organic carbon (SOC) through agricultural management could alter catchment-scale hydrology, using the Broye catchment in Western Switzerland as a representative case study. We specifically evaluated responses for four nested subcatchments where discharge observations were available. By representing SOC-driven changes in soil hydraulic properties in a large-scale hydrological model (mHM), we traced how increased water retention could affect evapotranspiration, subsurface runoff, and streamflow extremes. We applied five SOC scenarios varying in depth and magnitude to explore process sensitivity. While the direction and timing of SOC effects are credible, their magnitude remains uncertain due to limitations in pedotransfer functions and parameterization. At the catchment scale, the increase in SOC increased soil water content by 1.34 %–3.75 %, slightly increased evapotranspiration by 0.15 %–0.38 %, and marginally reduced dicharge by 0.27 %–0.7 %, depending on the applied SOC scenario. Effects were highly context-dependent: SOC-driven improvements in soil water retention tended to support higher evapotranspiration but reduced groundwater recharge and discharge, a clear trade-off. These shifts occasionally intensified low flows in warmer and drier subcatchments (Arbogne, Petit Glâne), while they could temporarily alleviate them in cooler and wetter areas (Broye subcatchment, Flon), especially under deeper SOC increases.</p>
      <p id="d2e6490">Our key findings are: <list list-type="bullet"><list-item>
      <p id="d2e6495">Even optimistic and substantial increases of SOC, and thus changes in hydraulic properties, lead to relatively modest impacts at the catchment scale.</p></list-item><list-item>
      <p id="d2e6499">The hydrological effects of SOC management depend strongly on local hydro-climatic conditions: the intended increase in plant-available water can reduce critical low-flow periods. However, it can also lead to unwanted ET increases and slightly reduce summer discharge.</p></list-item><list-item>
      <p id="d2e6503">Future work should focus on capturing vegetation and transpiration dynamics more accurately, including the interplay of crops with different growing seasons (winter vs. spring crops), to improve model realism.</p></list-item></list></p>
      <p id="d2e6506">Overall, our analysis emphasizes the need for a better understanding of the trade-offs and balances between agricultural practices aimed at increasing soil organic carbon (SOC) – including initiatives such as the 4 per mille and other soil carbon sequestration efforts – and their resulting impacts on catchment hydrological processes, ranging from soil moisture dynamics to groundwater recharge and hydrologic extremes.</p>
</sec>

      
      </body>
    <back><app-group>

<app id="App1.Ch1.S1">
  <label>Appendix A</label><title>Water balance for default and optimized parameter sets and default and optimized precipitation input data</title>

<table-wrap id="TA1"><label>Table A1</label><caption><p id="d2e6525">Mean annual water balance components (2016–2022) for four subcatchments under three model runs. Where (i) is the run with the default parameter set and the default RhiresD precipitation input data, (ii) is the default parameter set but with the adjusted precipitation input data (RhiresD+) and (iii) is the optimized parameter set and precipitation input data. <inline-formula><mml:math id="M403" display="inline"><mml:mrow><mml:msub><mml:mi>Q</mml:mi><mml:mi mathvariant="normal">obs</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> and <inline-formula><mml:math id="M404" display="inline"><mml:mrow><mml:msub><mml:mi>Q</mml:mi><mml:mi mathvariant="normal">sim</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> denote the observed and simulated discharge, <inline-formula><mml:math id="M405" display="inline"><mml:mi>P</mml:mi></mml:math></inline-formula> is precipitation and ET is simulated evapotranspiration. Values in mm yr<sup>−1</sup>.</p></caption><oasis:table frame="topbot"><oasis:tgroup cols="6">
     <oasis:colspec colnum="1" colname="col1" align="left"/>
     <oasis:colspec colnum="2" colname="col2" align="left"/>
     <oasis:colspec colnum="3" colname="col3" align="center"/>
     <oasis:colspec colnum="4" colname="col4" align="center"/>
     <oasis:colspec colnum="5" colname="col5" align="center"/>
     <oasis:colspec colnum="6" colname="col6" align="center"/>
     <oasis:thead>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1">Run</oasis:entry>
         <oasis:entry colname="col2">Subcatchment</oasis:entry>
         <oasis:entry colname="col3"><inline-formula><mml:math id="M407" display="inline"><mml:mrow><mml:msub><mml:mi>Q</mml:mi><mml:mi mathvariant="normal">obs</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col4"><inline-formula><mml:math id="M408" display="inline"><mml:mrow><mml:msub><mml:mi>Q</mml:mi><mml:mi mathvariant="normal">sim</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col5"><inline-formula><mml:math id="M409" display="inline"><mml:mi>P</mml:mi></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col6">ET</oasis:entry>
       </oasis:row>
     </oasis:thead>
     <oasis:tbody>
       <oasis:row>
         <oasis:entry colname="col1">(i) Default <inline-formula><mml:math id="M410" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula> RhiresD</oasis:entry>
         <oasis:entry colname="col2">Broye (Payerne)</oasis:entry>
         <oasis:entry colname="col3">504</oasis:entry>
         <oasis:entry colname="col4">509</oasis:entry>
         <oasis:entry colname="col5">1120</oasis:entry>
         <oasis:entry colname="col6">586</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"/>
         <oasis:entry colname="col2">Petit Glâne (Cugy)</oasis:entry>
         <oasis:entry colname="col3">298</oasis:entry>
         <oasis:entry colname="col4">385</oasis:entry>
         <oasis:entry colname="col5">917</oasis:entry>
         <oasis:entry colname="col6">566</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"/>
         <oasis:entry colname="col2">Arbogne (Avenches)</oasis:entry>
         <oasis:entry colname="col3">244</oasis:entry>
         <oasis:entry colname="col4">370</oasis:entry>
         <oasis:entry colname="col5">867</oasis:entry>
         <oasis:entry colname="col6">577</oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1"/>
         <oasis:entry colname="col2">Flon Aval (Oron)</oasis:entry>
         <oasis:entry colname="col3">527</oasis:entry>
         <oasis:entry colname="col4">654</oasis:entry>
         <oasis:entry colname="col5">1285</oasis:entry>
         <oasis:entry colname="col6">615</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">(ii) Default <inline-formula><mml:math id="M411" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula> RhiresD+</oasis:entry>
         <oasis:entry colname="col2">Broye (Payerne)</oasis:entry>
         <oasis:entry colname="col3">504</oasis:entry>
         <oasis:entry colname="col4">509</oasis:entry>
         <oasis:entry colname="col5">1120</oasis:entry>
         <oasis:entry colname="col6">586</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"/>
         <oasis:entry colname="col2">Petit Glâne (Cugy)</oasis:entry>
         <oasis:entry colname="col3">298</oasis:entry>
         <oasis:entry colname="col4">373</oasis:entry>
         <oasis:entry colname="col5">917</oasis:entry>
         <oasis:entry colname="col6">545</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"/>
         <oasis:entry colname="col2">Arbogne (Avenches)</oasis:entry>
         <oasis:entry colname="col3">244</oasis:entry>
         <oasis:entry colname="col4">285</oasis:entry>
         <oasis:entry colname="col5">867</oasis:entry>
         <oasis:entry colname="col6">554</oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1"/>
         <oasis:entry colname="col2">Flon Aval (Oron)</oasis:entry>
         <oasis:entry colname="col3">527</oasis:entry>
         <oasis:entry colname="col4">654</oasis:entry>
         <oasis:entry colname="col5">1285</oasis:entry>
         <oasis:entry colname="col6">615</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">(iii) Calibrated <inline-formula><mml:math id="M412" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula> RhiresD+</oasis:entry>
         <oasis:entry colname="col2">Broye (Payerne)</oasis:entry>
         <oasis:entry colname="col3">504</oasis:entry>
         <oasis:entry colname="col4">490</oasis:entry>
         <oasis:entry colname="col5">1120</oasis:entry>
         <oasis:entry colname="col6">612</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"/>
         <oasis:entry colname="col2">Petit Glâne  (Cugy)</oasis:entry>
         <oasis:entry colname="col3">298</oasis:entry>
         <oasis:entry colname="col4">324</oasis:entry>
         <oasis:entry colname="col5">917</oasis:entry>
         <oasis:entry colname="col6">598</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"/>
         <oasis:entry colname="col2">Arbogne (Avenches)</oasis:entry>
         <oasis:entry colname="col3">244</oasis:entry>
         <oasis:entry colname="col4">260</oasis:entry>
         <oasis:entry colname="col5">867</oasis:entry>
         <oasis:entry colname="col6">589</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"/>
         <oasis:entry colname="col2">Flon Aval (Oron)</oasis:entry>
         <oasis:entry colname="col3">527</oasis:entry>
         <oasis:entry colname="col4">644</oasis:entry>
         <oasis:entry colname="col5">1285</oasis:entry>
         <oasis:entry colname="col6">635</oasis:entry>
       </oasis:row>
     </oasis:tbody>
   </oasis:tgroup></oasis:table></table-wrap>

</app>

<app id="App1.Ch1.S2">
  <label>Appendix B</label><title>Model parameterization</title>
<sec id="App1.Ch1.S2.SS1">
  <label>B1</label><title>Hydraulic Parameter Estimation</title>
      <p id="d2e6931">The following equations are used to estimate van Genuchten parameters and other key soil hydraulic properties (assuming soil texture given in fractions [0–1]):

                <disp-formula specific-use="gather" content-type="numbered"><mml:math id="M413" display="block"><mml:mtable displaystyle="true"><mml:mlabeledtr id="App1.Ch1.S2.E8"><mml:mtd><mml:mtext>B1</mml:mtext></mml:mtd><mml:mtd><mml:mrow><mml:mstyle class="stylechange" displaystyle="true"/><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub><mml:mo>=</mml:mo><mml:msub><mml:mi>P</mml:mi><mml:mi mathvariant="normal">constant</mml:mi></mml:msub><mml:mo>+</mml:mo><mml:msub><mml:mi>P</mml:mi><mml:mi mathvariant="normal">clay</mml:mi></mml:msub><mml:mo>⋅</mml:mo><mml:msub><mml:mi>T</mml:mi><mml:mi mathvariant="normal">clay</mml:mi></mml:msub><mml:mo>+</mml:mo><mml:msub><mml:mi>P</mml:mi><mml:mi mathvariant="normal">BD</mml:mi></mml:msub><mml:mo>⋅</mml:mo><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi mathvariant="normal">b</mml:mi></mml:msub></mml:mrow></mml:mtd></mml:mlabeledtr><mml:mlabeledtr id="App1.Ch1.S2.E9"><mml:mtd><mml:mtext>B2</mml:mtext></mml:mtd><mml:mtd><mml:mrow><mml:mstyle displaystyle="true" class="stylechange"/><mml:mi>n</mml:mi><mml:mo>=</mml:mo><mml:msub><mml:mi>C</mml:mi><mml:mrow><mml:mi mathvariant="normal">vG</mml:mi><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msub><mml:mo>-</mml:mo><mml:msub><mml:mi>C</mml:mi><mml:mrow><mml:mi mathvariant="normal">vG</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:mrow></mml:msub><mml:mo>⋅</mml:mo><mml:msubsup><mml:mi>T</mml:mi><mml:mi mathvariant="normal">sand</mml:mi><mml:mrow><mml:msub><mml:mi>C</mml:mi><mml:mrow><mml:mi mathvariant="normal">vG</mml:mi><mml:mn mathvariant="normal">3</mml:mn></mml:mrow></mml:msub></mml:mrow></mml:msubsup><mml:mo>+</mml:mo><mml:msub><mml:mi>C</mml:mi><mml:mrow><mml:mi mathvariant="normal">vG</mml:mi><mml:mn mathvariant="normal">4</mml:mn></mml:mrow></mml:msub><mml:mo>⋅</mml:mo><mml:msubsup><mml:mi>T</mml:mi><mml:mi mathvariant="normal">clay</mml:mi><mml:mrow><mml:msub><mml:mi>C</mml:mi><mml:mrow><mml:mi mathvariant="normal">vG</mml:mi><mml:mn mathvariant="normal">5</mml:mn></mml:mrow></mml:msub></mml:mrow></mml:msubsup></mml:mrow></mml:mtd></mml:mlabeledtr><mml:mlabeledtr id="App1.Ch1.S2.E10"><mml:mtd><mml:mtext>B3</mml:mtext></mml:mtd><mml:mtd><mml:mrow><mml:mstyle class="stylechange" displaystyle="true"/><mml:mi>m</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mo>-</mml:mo><mml:mstyle displaystyle="true"><mml:mfrac style="display"><mml:mn mathvariant="normal">1</mml:mn><mml:mi>n</mml:mi></mml:mfrac></mml:mstyle></mml:mrow></mml:mtd></mml:mlabeledtr><mml:mlabeledtr id="App1.Ch1.S2.E11"><mml:mtd><mml:mtext>B4</mml:mtext></mml:mtd><mml:mtd><mml:mrow><mml:mstyle class="stylechange" displaystyle="true"/><mml:mi mathvariant="italic">α</mml:mi><mml:mo>=</mml:mo><mml:mi>exp⁡</mml:mi><mml:mo mathsize="1.1em">(</mml:mo><mml:msub><mml:mi>C</mml:mi><mml:mrow><mml:mi mathvariant="normal">vG</mml:mi><mml:mn mathvariant="normal">6</mml:mn></mml:mrow></mml:msub><mml:mo>+</mml:mo><mml:msub><mml:mi>C</mml:mi><mml:mrow><mml:mi mathvariant="normal">vG</mml:mi><mml:mn mathvariant="normal">7</mml:mn></mml:mrow></mml:msub><mml:mo>⋅</mml:mo><mml:msub><mml:mi>T</mml:mi><mml:mi mathvariant="normal">sand</mml:mi></mml:msub><mml:mo>+</mml:mo><mml:msub><mml:mi>C</mml:mi><mml:mrow><mml:mi mathvariant="normal">vG</mml:mi><mml:mn mathvariant="normal">8</mml:mn></mml:mrow></mml:msub><mml:mo>⋅</mml:mo><mml:msub><mml:mi>T</mml:mi><mml:mi mathvariant="normal">clay</mml:mi></mml:msub><mml:mo>-</mml:mo><mml:msub><mml:mi>C</mml:mi><mml:mrow><mml:mi mathvariant="normal">vG</mml:mi><mml:mn mathvariant="normal">9</mml:mn></mml:mrow></mml:msub><mml:mo>⋅</mml:mo><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi mathvariant="normal">b</mml:mi></mml:msub><mml:mo mathsize="1.1em">)</mml:mo></mml:mrow></mml:mtd></mml:mlabeledtr><mml:mlabeledtr id="App1.Ch1.S2.E12"><mml:mtd><mml:mtext>B5</mml:mtext></mml:mtd><mml:mtd><mml:mrow><mml:mstyle class="stylechange" displaystyle="true"/><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">FC</mml:mi></mml:msub><mml:mo>=</mml:mo><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub><mml:mo>⋅</mml:mo><mml:mi>exp⁡</mml:mi><mml:mfenced open="(" close=")"><mml:mrow><mml:msub><mml:mi>C</mml:mi><mml:mrow><mml:mi mathvariant="normal">FC</mml:mi><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msub><mml:mo>⋅</mml:mo><mml:mo mathsize="1.1em">(</mml:mo><mml:msub><mml:mi>C</mml:mi><mml:mrow><mml:mi mathvariant="normal">FC</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:mrow></mml:msub><mml:mo>+</mml:mo><mml:msub><mml:mi>log⁡</mml:mi><mml:mn mathvariant="normal">10</mml:mn></mml:msub><mml:mo>(</mml:mo><mml:msub><mml:mi>K</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub><mml:mo>)</mml:mo><mml:mo mathsize="1.1em">)</mml:mo><mml:mo>⋅</mml:mo><mml:mi>log⁡</mml:mi><mml:mo>(</mml:mo><mml:mi>n</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:mfenced></mml:mrow></mml:mtd></mml:mlabeledtr><mml:mlabeledtr id="App1.Ch1.S2.E13"><mml:mtd><mml:mtext>B6</mml:mtext></mml:mtd><mml:mtd><mml:mrow><mml:mstyle displaystyle="true" class="stylechange"/><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">PWP</mml:mi></mml:msub><mml:mo>=</mml:mo><mml:mstyle displaystyle="true"><mml:mfrac style="display"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow><mml:mrow><mml:mi>exp⁡</mml:mi><mml:mfenced open="(" close=")"><mml:mrow><mml:mi>m</mml:mi><mml:mspace linebreak="nobreak" width="0.33em"/><mml:mo>⋅</mml:mo><mml:mi>log⁡</mml:mi><mml:mfenced open="(" close=")"><mml:mrow><mml:msub><mml:mi>C</mml:mi><mml:mi mathvariant="normal">PWPc</mml:mi></mml:msub><mml:mo>+</mml:mo><mml:mi>exp⁡</mml:mi><mml:mfenced close=")" open="("><mml:mrow><mml:mi>n</mml:mi><mml:mo>⋅</mml:mo><mml:mi>log⁡</mml:mi><mml:mfenced open="(" close=")"><mml:mrow><mml:mi mathvariant="italic">α</mml:mi><mml:mo>⋅</mml:mo><mml:msub><mml:mi>C</mml:mi><mml:mi mathvariant="normal">PWPh</mml:mi></mml:msub></mml:mrow></mml:mfenced></mml:mrow></mml:mfenced></mml:mrow></mml:mfenced></mml:mrow></mml:mfenced></mml:mrow></mml:mfrac></mml:mstyle></mml:mrow></mml:mtd></mml:mlabeledtr></mml:mtable></mml:math></disp-formula>

          All constant and parameter values are listed in the Sect. S4. Equation for <inline-formula><mml:math id="M414" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">FC</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> depends on paper by <xref ref-type="bibr" rid="bib1.bibx123" id="text.170"/>, that calculates <inline-formula><mml:math id="M415" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">FC</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> in dependence of <inline-formula><mml:math id="M416" display="inline"><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, decreasing <inline-formula><mml:math id="M417" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">FC</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> with increasing <inline-formula><mml:math id="M418" display="inline"><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>. Changes in SOC and bulk density (<inline-formula><mml:math id="M419" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi mathvariant="normal">b</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>) would propagate to other soil hydraulic parameters, as evident in Eqs. (<xref ref-type="disp-formula" rid="App1.Ch1.S2.E8"/>) to (<xref ref-type="disp-formula" rid="App1.Ch1.S2.E13"/>). These would affect the estimation of <inline-formula><mml:math id="M420" display="inline"><mml:mi mathvariant="italic">θ</mml:mi></mml:math></inline-formula>, which is initialized to 0.25 and then​​​​​​​​​​​​​​​​  updated at each timestep via:

                <disp-formula specific-use="gather" content-type="numbered"><mml:math id="M421" display="block"><mml:mtable displaystyle="true"><mml:mlabeledtr id="App1.Ch1.S2.E14"><mml:mtd><mml:mtext>B7</mml:mtext></mml:mtd><mml:mtd><mml:mrow><mml:mstyle displaystyle="true" class="stylechange"/><mml:mi>I</mml:mi><mml:mo>=</mml:mo><mml:mfenced open="{" close=""><mml:mtable columnspacing="1em" class="cases" rowspacing="0.2ex" columnalign="left left" framespacing="0em"><mml:mtr><mml:mtd><mml:mrow><mml:msub><mml:mi>P</mml:mi><mml:mi mathvariant="normal">effective</mml:mi></mml:msub><mml:mo>,</mml:mo></mml:mrow></mml:mtd><mml:mtd><mml:mrow><mml:mi mathvariant="normal">if</mml:mi><mml:mspace width="0.25em" linebreak="nobreak"/><mml:mi mathvariant="italic">θ</mml:mi><mml:mo>&gt;</mml:mo><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:mtd></mml:mtr><mml:mtr><mml:mtd><mml:mrow><mml:msub><mml:mi>P</mml:mi><mml:mi mathvariant="normal">effective</mml:mi></mml:msub><mml:mo>+</mml:mo><mml:mo>(</mml:mo><mml:mi mathvariant="italic">θ</mml:mi><mml:mo>-</mml:mo><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub><mml:mo>)</mml:mo><mml:mo>,</mml:mo></mml:mrow></mml:mtd><mml:mtd><mml:mrow><mml:mi mathvariant="normal">if</mml:mi><mml:mspace linebreak="nobreak" width="0.25em"/><mml:mi mathvariant="italic">θ</mml:mi><mml:mo>+</mml:mo><mml:msub><mml:mi>x</mml:mi><mml:mi mathvariant="normal">tmp</mml:mi></mml:msub><mml:mo>&gt;</mml:mo><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:mtd></mml:mtr><mml:mtr><mml:mtd><mml:mrow><mml:msub><mml:mi>P</mml:mi><mml:mi mathvariant="normal">effective</mml:mi></mml:msub><mml:mo>-</mml:mo><mml:msub><mml:mi>x</mml:mi><mml:mi mathvariant="normal">tmp</mml:mi></mml:msub><mml:mo>,</mml:mo></mml:mrow></mml:mtd><mml:mtd><mml:mi mathvariant="normal">otherwise</mml:mi></mml:mtd></mml:mtr></mml:mtable></mml:mfenced></mml:mrow></mml:mtd></mml:mlabeledtr><mml:mlabeledtr id="App1.Ch1.S2.E15"><mml:mtd><mml:mtext>B8</mml:mtext></mml:mtd><mml:mtd><mml:mrow><mml:mstyle displaystyle="true" class="stylechange"/><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">new</mml:mi></mml:msub><mml:mo>=</mml:mo><mml:mfenced open="{" close=""><mml:mtable class="cases" rowspacing="0.2ex" columnspacing="1em" columnalign="left left" framespacing="0em"><mml:mtr><mml:mtd><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub><mml:mo>,</mml:mo></mml:mrow></mml:mtd><mml:mtd><mml:mrow><mml:mi mathvariant="normal">if</mml:mi><mml:mspace width="0.25em" linebreak="nobreak"/><mml:mi mathvariant="italic">θ</mml:mi><mml:mo>+</mml:mo><mml:msub><mml:mi>x</mml:mi><mml:mi mathvariant="normal">tmp</mml:mi></mml:msub><mml:mo>&gt;</mml:mo><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:mtd></mml:mtr><mml:mtr><mml:mtd><mml:mrow><mml:mi mathvariant="italic">θ</mml:mi><mml:mo>+</mml:mo><mml:msub><mml:mi>x</mml:mi><mml:mi mathvariant="normal">tmp</mml:mi></mml:msub><mml:mo>,</mml:mo></mml:mrow></mml:mtd><mml:mtd><mml:mi mathvariant="normal">otherwise</mml:mi></mml:mtd></mml:mtr></mml:mtable></mml:mfenced></mml:mrow></mml:mtd></mml:mlabeledtr><mml:mlabeledtr id="App1.Ch1.S2.E16"><mml:mtd><mml:mtext>B9</mml:mtext></mml:mtd><mml:mtd><mml:mrow><mml:mstyle class="stylechange" displaystyle="true"/><mml:msub><mml:mi mathvariant="normal">f</mml:mi><mml:mi mathvariant="normal">runoff</mml:mi></mml:msub><mml:mo>=</mml:mo><mml:mfenced open="{" close=""><mml:mtable rowspacing="0.2ex" class="cases" columnspacing="1em" columnalign="left left" framespacing="0em"><mml:mtr><mml:mtd><mml:mrow><mml:mi>exp⁡</mml:mi><mml:mo>(</mml:mo><mml:msub><mml:mi>e</mml:mi><mml:mrow><mml:mi mathvariant="normal">soil</mml:mi><mml:mi mathvariant="normal">_</mml:mi><mml:mi mathvariant="normal">moisture</mml:mi></mml:mrow></mml:msub><mml:mi>log⁡</mml:mi><mml:mo>(</mml:mo><mml:mstyle displaystyle="false"><mml:mfrac style="text"><mml:mi mathvariant="italic">θ</mml:mi><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">sat</mml:mi></mml:msub></mml:mrow></mml:mfrac></mml:mstyle><mml:mo>)</mml:mo><mml:mo>)</mml:mo><mml:mo>,</mml:mo></mml:mrow></mml:mtd><mml:mtd><mml:mrow><mml:mi mathvariant="italic">θ</mml:mi><mml:mo>&gt;</mml:mo><mml:mn mathvariant="normal">0</mml:mn></mml:mrow></mml:mtd></mml:mtr><mml:mtr><mml:mtd><mml:mrow><mml:mn mathvariant="normal">0</mml:mn><mml:mo>,</mml:mo></mml:mrow></mml:mtd><mml:mtd><mml:mi mathvariant="normal">otherwise</mml:mi></mml:mtd></mml:mtr></mml:mtable></mml:mfenced></mml:mrow></mml:mtd></mml:mlabeledtr><mml:mlabeledtr id="App1.Ch1.S2.E17"><mml:mtd><mml:mtext>B10</mml:mtext></mml:mtd><mml:mtd><mml:mrow><mml:mstyle class="stylechange" displaystyle="true"/><mml:msub><mml:mi>x</mml:mi><mml:mi mathvariant="normal">tmp</mml:mi></mml:msub><mml:mo>=</mml:mo><mml:msub><mml:mi>P</mml:mi><mml:mi mathvariant="normal">effective</mml:mi></mml:msub><mml:mfenced open="(" close=")"><mml:mrow><mml:mn mathvariant="normal">1</mml:mn><mml:mo>-</mml:mo><mml:msub><mml:mi mathvariant="normal">f</mml:mi><mml:mi mathvariant="normal">runoff</mml:mi></mml:msub></mml:mrow></mml:mfenced></mml:mrow></mml:mtd></mml:mlabeledtr></mml:mtable></mml:math></disp-formula>

          where <inline-formula><mml:math id="M422" display="inline"><mml:mrow><mml:msub><mml:mi>P</mml:mi><mml:mi mathvariant="normal">effective</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> is either incoming precipitation or Infiltration (<inline-formula><mml:math id="M423" display="inline"><mml:mi>I</mml:mi></mml:math></inline-formula>) from the above soil layer. The change in <inline-formula><mml:math id="M424" display="inline"><mml:mi mathvariant="italic">θ</mml:mi></mml:math></inline-formula> then again propagates to the root zone soil moisture storage (<inline-formula><mml:math id="M425" display="inline"><mml:mrow><mml:msub><mml:mi>X</mml:mi><mml:mn mathvariant="normal">3</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> in Fig. <xref ref-type="fig" rid="F1"/>):

            <disp-formula id="App1.Ch1.S2.E18" content-type="numbered"><label>B11</label><mml:math id="M426" display="block"><mml:mrow><mml:msub><mml:mi>X</mml:mi><mml:mn mathvariant="normal">3</mml:mn></mml:msub><mml:mo>=</mml:mo><mml:msup><mml:mi>I</mml:mi><mml:mrow><mml:mo>(</mml:mo><mml:mi>k</mml:mi><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mo>)</mml:mo></mml:mrow></mml:msup><mml:mo>-</mml:mo><mml:msup><mml:mi mathvariant="normal">ET</mml:mi><mml:mi>k</mml:mi></mml:msup><mml:mo>-</mml:mo><mml:mfenced close=")" open="("><mml:mrow><mml:msup><mml:mi>I</mml:mi><mml:mrow><mml:mo>(</mml:mo><mml:mi>k</mml:mi><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mo>)</mml:mo></mml:mrow></mml:msup><mml:mo>-</mml:mo><mml:mfenced open="(" close=")"><mml:mrow><mml:msup><mml:mi>I</mml:mi><mml:mrow><mml:mo>(</mml:mo><mml:mi>k</mml:mi><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mo>)</mml:mo></mml:mrow></mml:msup><mml:mfenced open="(" close=")"><mml:mrow><mml:mn mathvariant="normal">1</mml:mn><mml:mo>-</mml:mo><mml:msub><mml:mi mathvariant="normal">f</mml:mi><mml:mi mathvariant="normal">runoff</mml:mi></mml:msub></mml:mrow></mml:mfenced></mml:mrow></mml:mfenced></mml:mrow></mml:mfenced></mml:mrow></mml:math></disp-formula>

          where <inline-formula><mml:math id="M427" display="inline"><mml:mi>k</mml:mi></mml:math></inline-formula> is the soil layer and <inline-formula><mml:math id="M428" display="inline"><mml:mi>I</mml:mi></mml:math></inline-formula> is the Infiltration coming from the layer above and <inline-formula><mml:math id="M429" display="inline"><mml:mrow><mml:msub><mml:mi>e</mml:mi><mml:mrow><mml:mi mathvariant="normal">soil</mml:mi><mml:mi mathvariant="normal">_</mml:mi><mml:mi mathvariant="normal">moisture</mml:mi></mml:mrow></mml:msub></mml:mrow></mml:math></inline-formula> is being calibrated.</p>
</sec>
<sec id="App1.Ch1.S2.SS2">
  <label>B2</label><title>Field Capacity Normalization and Soil stress factor calculation</title>
      <p id="d2e7821">Not only do the changes in soil hydraulic properties affect soil moisture, but the soil moisture also governs how much water can evapotranspire from each layer:

            <disp-formula id="App1.Ch1.S2.E19" content-type="numbered"><label>B12</label><mml:math id="M430" display="block"><mml:mrow><mml:mover accent="true"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">FC</mml:mi></mml:msub></mml:mrow><mml:mo mathvariant="normal">‾</mml:mo></mml:mover><mml:mo>=</mml:mo><mml:mstyle displaystyle="true"><mml:mfrac style="display"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">FC</mml:mi></mml:msub><mml:mo>-</mml:mo><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">min</mml:mi></mml:msub></mml:mrow><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">max</mml:mi></mml:msub><mml:mo>-</mml:mo><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">min</mml:mi></mml:msub></mml:mrow></mml:mfrac></mml:mstyle></mml:mrow></mml:math></disp-formula>

          where <inline-formula><mml:math id="M431" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">max</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> is <inline-formula><mml:math id="M432" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">global</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> + <inline-formula><mml:math id="M433" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">min</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, and:

            <disp-formula id="App1.Ch1.S2.E20" content-type="numbered"><label>B13</label><mml:math id="M434" display="block"><mml:mrow><mml:msub><mml:mi>R</mml:mi><mml:mi mathvariant="normal">CoeffFC</mml:mi></mml:msub><mml:mo>=</mml:mo><mml:mover accent="true"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">FC</mml:mi></mml:msub></mml:mrow><mml:mo mathvariant="normal">‾</mml:mo></mml:mover><mml:mo>⋅</mml:mo><mml:msub><mml:mi>f</mml:mi><mml:mi mathvariant="normal">clay</mml:mi></mml:msub><mml:mo>+</mml:mo><mml:mo>(</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mo>-</mml:mo><mml:mover accent="true"><mml:mrow><mml:msub><mml:mi mathvariant="italic">θ</mml:mi><mml:mi mathvariant="normal">FC</mml:mi></mml:msub></mml:mrow><mml:mo mathvariant="normal">‾</mml:mo></mml:mover><mml:mo>)</mml:mo><mml:mo>⋅</mml:mo><mml:msub><mml:mi>f</mml:mi><mml:mi mathvariant="normal">sand</mml:mi></mml:msub></mml:mrow></mml:math></disp-formula></p>
</sec>
</app>

<app id="App1.Ch1.S3">
  <label>Appendix C</label><title>Model outputs and evaluation</title>
<sec id="App1.Ch1.S3.SS1">
  <label>C1</label><title>Observed and simulated soil moisture</title>

      <fig id="FC1"><label>Figure C1</label><caption><p id="d2e7975">Observed and simulated (mHM) timeseries of volumetric water content at the SwissSMEX grassland site near Payerne. Simulations represent three soil layers, while observations are point-scale: layer 1 (5, 10, 15 cm, integrated), layer 2 (50 cm), and layer 3 (80 cm).</p></caption>
          
          <graphic xlink:href="https://hess.copernicus.org/articles/30/2879/2026/hess-30-2879-2026-f13.png"/>

        </fig>


</sec>
<sec id="App1.Ch1.S3.SS2">
  <label>C2</label><title>Absolute differences in discharge</title>

      <fig id="FC2"><label>Figure C2</label><caption><p id="d2e7998">Absolute difference in discharge between the base and each SOC increase scenarios.</p></caption>
          
          <graphic xlink:href="https://hess.copernicus.org/articles/30/2879/2026/hess-30-2879-2026-f14.png"/>

        </fig>


</sec>
<sec id="App1.Ch1.S3.SS3">
  <label>C3</label><title>Timing of simulated low flow periods</title>

      <fig id="FC3"><label>Figure C3</label><caption><p id="d2e8021">Observed and simulated low flow days for all subcatchments.</p></caption>
          
          <graphic xlink:href="https://hess.copernicus.org/articles/30/2879/2026/hess-30-2879-2026-f15.png"/>

        </fig>

</sec>
<sec id="App1.Ch1.S3.SS4">
  <label>C4</label><title>Evaluation of simulated discharge dynamics</title>

<table-wrap id="TC1"><label>Table C1</label><caption><p id="d2e8044">Metrics for peak flow and low flow fit: <inline-formula><mml:math id="M435" display="inline"><mml:mrow><mml:msub><mml:mi>Q</mml:mi><mml:mn mathvariant="normal">95</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> and <inline-formula><mml:math id="M436" display="inline"><mml:mrow><mml:msub><mml:mi>Q</mml:mi><mml:mn mathvariant="normal">5</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> denote high- and low-flow percentiles; Peak_bias and Low_bias are percent biases (%).</p></caption><oasis:table frame="topbot"><oasis:tgroup cols="9">
     <oasis:colspec colnum="1" colname="col1" align="left"/>
     <oasis:colspec colnum="2" colname="col2" align="right"/>
     <oasis:colspec colnum="3" colname="col3" align="right"/>
     <oasis:colspec colnum="4" colname="col4" align="right"/>
     <oasis:colspec colnum="5" colname="col5" align="right"/>
     <oasis:colspec colnum="6" colname="col6" align="right"/>
     <oasis:colspec colnum="7" colname="col7" align="right"/>
     <oasis:colspec colnum="8" colname="col8" align="right"/>
     <oasis:colspec colnum="9" colname="col9" align="left"/>
     <oasis:thead>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1">Station</oasis:entry>
         <oasis:entry colname="col2">KGE</oasis:entry>
         <oasis:entry colname="col3"><inline-formula><mml:math id="M437" display="inline"><mml:mrow><mml:msub><mml:mi>Q</mml:mi><mml:mrow><mml:msub><mml:mn mathvariant="normal">95</mml:mn><mml:mi mathvariant="normal">obs</mml:mi></mml:msub></mml:mrow></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col4"><inline-formula><mml:math id="M438" display="inline"><mml:mrow><mml:msub><mml:mi>Q</mml:mi><mml:mrow><mml:msub><mml:mn mathvariant="normal">95</mml:mn><mml:mi mathvariant="normal">sim</mml:mi></mml:msub></mml:mrow></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col5"><inline-formula><mml:math id="M439" display="inline"><mml:mrow><mml:msub><mml:mi>Q</mml:mi><mml:mrow><mml:msub><mml:mn mathvariant="normal">5</mml:mn><mml:mi mathvariant="normal">obs</mml:mi></mml:msub></mml:mrow></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col6"><inline-formula><mml:math id="M440" display="inline"><mml:mrow><mml:msub><mml:mi>Q</mml:mi><mml:mrow><mml:msub><mml:mn mathvariant="normal">5</mml:mn><mml:mi mathvariant="normal">sim</mml:mi></mml:msub></mml:mrow></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col7">Peak<sub>bias</sub></oasis:entry>
         <oasis:entry colname="col8">Low<sub>bias</sub></oasis:entry>
         <oasis:entry colname="col9"/>
       </oasis:row>
     </oasis:thead>
     <oasis:tbody>
       <oasis:row>
         <oasis:entry colname="col1">Petit Glâne</oasis:entry>
         <oasis:entry colname="col2">0.86</oasis:entry>
         <oasis:entry colname="col3">2.63</oasis:entry>
         <oasis:entry colname="col4">2.66</oasis:entry>
         <oasis:entry colname="col5">0.17</oasis:entry>
         <oasis:entry colname="col6">0.20</oasis:entry>
         <oasis:entry colname="col7">1.24</oasis:entry>
         <oasis:entry colname="col8">18.09</oasis:entry>
         <oasis:entry colname="col9"/>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Arbogne</oasis:entry>
         <oasis:entry colname="col2">0.83</oasis:entry>
         <oasis:entry colname="col3">1.77</oasis:entry>
         <oasis:entry colname="col4">1.76</oasis:entry>
         <oasis:entry colname="col5">0.20</oasis:entry>
         <oasis:entry colname="col6">0.15</oasis:entry>
         <oasis:entry colname="col7"><inline-formula><mml:math id="M443" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.08</oasis:entry>
         <oasis:entry colname="col8"><inline-formula><mml:math id="M444" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>22.51</oasis:entry>
         <oasis:entry colname="col9"/>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Flon</oasis:entry>
         <oasis:entry colname="col2">0.91</oasis:entry>
         <oasis:entry colname="col3">1.18</oasis:entry>
         <oasis:entry colname="col4">1.22</oasis:entry>
         <oasis:entry colname="col5">0.02</oasis:entry>
         <oasis:entry colname="col6">0.02</oasis:entry>
         <oasis:entry colname="col7">3.79</oasis:entry>
         <oasis:entry colname="col8">43.23</oasis:entry>
         <oasis:entry colname="col9"/>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Broye</oasis:entry>
         <oasis:entry colname="col2">0.91</oasis:entry>
         <oasis:entry colname="col3">21.02</oasis:entry>
         <oasis:entry colname="col4">22.49</oasis:entry>
         <oasis:entry colname="col5">0.87</oasis:entry>
         <oasis:entry colname="col6">0.94</oasis:entry>
         <oasis:entry colname="col7">6.98</oasis:entry>
         <oasis:entry colname="col8">8.57</oasis:entry>
         <oasis:entry colname="col9"/>
       </oasis:row>
     </oasis:tbody>
   </oasis:tgroup></oasis:table></table-wrap>


</sec>
<sec id="App1.Ch1.S3.SS5">
  <label>C5</label><title>Maps of key variables and fluxes</title>

      <fig id="FC4"><label>Figure C4</label><caption><p id="d2e8343">Monthly and annually aggregated spatial patterns of key fluxes. <inline-formula><mml:math id="M445" display="inline"><mml:mrow><mml:mi>Q</mml:mi><mml:mo>=</mml:mo></mml:mrow></mml:math></inline-formula> discharge, <inline-formula><mml:math id="M446" display="inline"><mml:mrow><mml:mi>P</mml:mi><mml:mo>=</mml:mo></mml:mrow></mml:math></inline-formula> precipitation, <inline-formula><mml:math id="M447" display="inline"><mml:mi>T</mml:mi></mml:math></inline-formula> mean <inline-formula><mml:math id="M448" display="inline"><mml:mo>=</mml:mo></mml:math></inline-formula> average temperature, ET <inline-formula><mml:math id="M449" display="inline"><mml:mo>/</mml:mo></mml:math></inline-formula> PET <inline-formula><mml:math id="M450" display="inline"><mml:mo>=</mml:mo></mml:math></inline-formula> ratio of actual to potential evapotranspiration, ET <inline-formula><mml:math id="M451" display="inline"><mml:mo>/</mml:mo></mml:math></inline-formula> <inline-formula><mml:math id="M452" display="inline"><mml:mi>P</mml:mi></mml:math></inline-formula> <inline-formula><mml:math id="M453" display="inline"><mml:mo>=</mml:mo></mml:math></inline-formula> ration of actual evapotranspiration to precipitation. <inline-formula><mml:math id="M454" display="inline"><mml:mrow><mml:msub><mml:mi>Q</mml:mi><mml:mi mathvariant="normal">diff</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M455" display="inline"><mml:mo>=</mml:mo></mml:math></inline-formula> absolute difference in discharge between base and example SOC increase scenario (MedC_LowC).</p></caption>
          
          <graphic xlink:href="https://hess.copernicus.org/articles/30/2879/2026/hess-30-2879-2026-f16.png"/>

        </fig>

</sec>
<sec id="App1.Ch1.S3.SS6">
  <label>C6</label><title>Impacts on peak flows (absolute differences)</title>

      <fig id="FC5"><label>Figure C5</label><caption><p id="d2e8452">Absolute difference in discharge for peak flow events for SOC scenarios vs. the base scenario. “<inline-formula><mml:math id="M456" display="inline"><mml:mi>Q</mml:mi></mml:math></inline-formula> base” denotes the absolute discharge value for each event.</p></caption>
          
          <graphic xlink:href="https://hess.copernicus.org/articles/30/2879/2026/hess-30-2879-2026-f17.png"/>

        </fig>


</sec>
</app>

<app id="App1.Ch1.S4">
  <label>Appendix D</label><title>Number of low flow days for different optimization runs</title>

      <fig id="FD1"><label>Figure D1</label><caption><p id="d2e8483">Difference in number of low flow days for base vs. SOC scenario (example scenario MedC_lowC) for 6 different optimization runs with the same setting.</p></caption>
        
        <graphic xlink:href="https://hess.copernicus.org/articles/30/2879/2026/hess-30-2879-2026-f18.png"/>

      </fig>

</app>
  </app-group><notes notes-type="codeavailability"><title>Code availability</title>

      <p id="d2e8498">Scripts to pre- and postprocess and visualize mHM input and output data <ext-link xlink:href="https://doi.org/10.5281/zenodo.17515165" ext-link-type="DOI">10.5281/zenodo.17515165</ext-link> <xref ref-type="bibr" rid="bib1.bibx48" id="paren.171"/>. The mHM source code is available on the developers GitHub <uri>https://github.com/mhm-ufz/mhm</uri> (last access: 10 May 2026;  <ext-link xlink:href="https://doi.org/10.5281/zenodo.1069202" ext-link-type="DOI">10.5281/zenodo.1069202</ext-link>, <xref ref-type="bibr" rid="bib1.bibx99" id="altparen.172"/>).</p>
  </notes><notes notes-type="dataavailability"><title>Data availability</title>

      <p id="d2e8519">The adapted precipitation timeseries (explained in further detail in Sect. <xref ref-type="sec" rid="Ch1.S2.SS5"/> and in the Supplement) is available here: <ext-link xlink:href="https://doi.org/10.5281/zenodo.17243147" ext-link-type="DOI">10.5281/zenodo.17243147</ext-link> <xref ref-type="bibr" rid="bib1.bibx47" id="paren.173"/>.</p>
  </notes><app-group>
        <supplementary-material position="anchor"><p id="d2e8530">The supplement related to this article is available online at <inline-supplementary-material xlink:href="https://doi.org/10.5194/hess-30-2879-2026-supplement" xlink:title="pdf">https://doi.org/10.5194/hess-30-2879-2026-supplement</inline-supplementary-material>.</p></supplementary-material>
        </app-group><notes notes-type="authorcontribution"><title>Author contributions</title>

      <p id="d2e8539">MH, BS, AH, PH and RK contributed to conceptualization; MH, BS, AH, PH and RK to methodology; MH, SL PH and RK to software; MH to validation; MH to formal analysis; MH to investigation; PH, BS and RK to resources; MH, PH and SL to data curation; MH, PH and SL to writing – original draft; MH, BS, AH PH and RK to writing – review &amp; editing; MH to visualization; AH, BS and RK to supervision.</p>
  </notes><notes notes-type="competinginterests"><title>Competing interests</title>

      <p id="d2e8548">At least one of the (co-)authors is a member of the editorial board of <italic>Hydrology and Earth System Sciences</italic>. The peer-review process was guided by an independent editor, and the authors also have no other competing interests to declare.</p>
  </notes><notes notes-type="disclaimer"><title>Disclaimer</title>

      <p id="d2e8558">Publisher's note: Copernicus Publications remains neutral with regard to jurisdictional claims made in the text, published maps, institutional affiliations, or any other geographical representation in this paper. The authors bear the ultimate responsibility for providing appropriate place names. Views expressed in the text are those of the authors and do not necessarily reflect the views of the publisher.</p>
  </notes><ack><title>Acknowledgements</title><p id="d2e8564">Calculations were performed on UBELIX (<uri>https://www.id.unibe.ch/hpc</uri>, last access: 10 May 2026), the HPC cluster at the University of Bern. We thank Christoph Raible and Natalie Ceperley for stimulating discussions on the manuscript. We thank Pallav Kumar Shrestha for a pre-release version of mHM including the SCC module,  described in the Methods section. OpenAI's ChatGPT (GPT-5) was used as a support tool for code development and for improving clarity and conciseness of the text. All content was reviewed, edited, and verified by the author, who assumes full responsibility for the work. The software BioRender was used in the creation of figures.</p></ack><notes notes-type="reviewstatement"><title>Review statement</title>

      <p id="d2e8572">This paper was edited by Julia Knapp and reviewed by Zhaoyang Luo and two anonymous referees.</p>
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